This is B-roll video of POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022.
Anthony M Pagano, Ph.D.
Ecophysiology, spatial ecology, and behavior of large mammals
Professional Experience
2022 - Present Research Wildlife Biologist USGS Alaska Science Center, Anchorage, Alaska
2021 - 2022 Post-doctoral Researcher Washington State University, Pullman, Washington
2018 - 2021 Post-doctoral Researcher Institute for Conservation Research, San Diego Zoo Wildlife Alliance, Escondido, California
2008 - 2018 Wildlife Biologist USGS Alaska Science Center, Anchorage, Alaska
Education and Certifications
Ph.D. 2018 University of California, Santa Cruz Ecology and Evolutionary Biology
M.S. 2007 University of Minnesota, St. Paul, MN Wildlife Conservation
B.A. 2002 Northeastern University, Boston, MA Biology and History
Science and Products
Polar Bear Research
Distribution and Movements of Polar Bears
Health and Energetics of Polar Bears
Metabolic Rate, Body Composition, and Blood Biochemistry Data from Polar Bears (Ursus maritimus) on Land, Western Hudson Bay, Canada, 2019-2022
Polar Bear Continuous Time-Correlated Random Walk (CTCRW) Location Data Derived from Satellite Location Data, Chukchi and Beaufort Seas, July-November 1985-2017
Polar Bear Continuous Time-Correlated Random Walk (CTCRW) Location Data Derived from Satellite Location Data, Southern Beaufort Sea, 1986-2016
Serum Urea and Creatinine Levels of Spring-Caught Polar Bears (Ursus maritimus) in the Southern Beaufort and Chukchi Seas
Satellite Location and Tri-axial Accelerometer Data from Adult Female Polar Bears (Ursus maritimus) in the Southern Beaufort Sea, April-October 2014
Locations Collected 1985-2015 from Female Polar Bears (Ursus maritimus) with Dependent Young Instrumented in the Southern Beaufort Sea with Satellite-linked Transmitters by the USGS
Bioelectrical Impedance, Deuterium Dilution, Body Mass, and Morphological Measures of Southern Beaufort Sea Female Polar Bears, Spring 2014-2016
Metabolic Rate, Body Composition, Foraging Success, Behavior, and GPS Locations of Female Polar Bears (Ursus maritimus), Beaufort Sea, Spring, 2014-2016 and Resting Energetics of an Adult Female Polar Bear
Energetic Costs of Locomotion in Bears
Measures of oxygen consumption and stroke frequency of a captive subadult polar bear (Ursus maritimus) while resting in water and swimming and diving in a metabolic water flume, Oregon Zoo, 2017
This is B-roll video of POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022.
An adult male polar bear with a GPS-enabled video camera collar on land in Wapusk National Park, Canada as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
An adult male polar bear with a GPS-enabled video camera collar on land in Wapusk National Park, Canada as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with another bear while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with another bear while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear walking along the coast in Wapusk National Park, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear walking along the coast in Wapusk National Park, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while on land near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while on land near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Ursids evolved dietary diversity without major alterations in metabolic rates
Polar bear energetic and behavioral strategies on land with implications for surviving the ice-free period
Declining Arctic sea ice is increasing polar bear land use. Polar bears on land are thought to minimize activity to conserve energy. Here, we measure the daily energy expenditure (DEE), diet, behavior, movement, and body composition changes of 20 different polar bears on land over 19–23 days from August to September (2019–2022) in Manitoba, Canada. Polar bears on land exhibited a 5.2-fold range in
Incremental evolution of modeling a prognosis for polar bears in a rapidly changing Arctic
A body composition model with multiple storage compartments for polar bears (Ursus maritimus)
Observed and forecasted changes in land use by polar bears in the Beaufort and Chukchi Seas, 1985–2040
Effects of sea ice decline and summer land use on polar bear home range size in the Beaufort Sea
Human-polar bear interactions
Polar bear foraging behavior
Identifying reliable indicators of fitness in polar bears
The seasonal energetic landscape of an apex marine carnivore, the polar bear
Overhauling ocean spatial planning to improve marine megafauna conservation
Energetic costs of aquatic locomotion in a subadult polar bear
Non-USGS Publications**
Cutting, N. Nicassio-Hiskey, A. Hash, and T.M. Williams. 2018. Energetic costs of
locomotion in bears: Is plantigrade locomotion energetically economical? Journal of
Experimental Biology vol. 221 no. 12 p.1-9. doi: 10.1242/jeb.175372
Costa, M.A. Owen, and T.M. Williams. 2018. High-energy, high-fat lifestyle challenges an
Arctic apex predator, the polar bear. Science vol. 359 no. 6375 p. 568-572. doi: 10.1126/science.aan8677
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
Science and Products
- Science
Polar Bear Research
Polar bears ( Ursus maritimus ) are one of 4 marine mammal species managed by the U.S. Department of Interior. The USGS Alaska Science Center leads long–term research on polar bears to inform local, state, national and international policy makers regarding conservation of the species and its habitat. Our studies, ongoing since 1985, are focused on population dynamics, health and energetics...Distribution and Movements of Polar Bears
Polar bears are tied to the sea ice for nearly all of their life cycle functions. Most important of these is foraging, or access to food. Polar bears almost exclusively eat seals, and they are equally as dependent upon the sea for their nutrition as are seals, whales, and other aquatic mammals. Polar bears are not aquatic, however, and their only access to the seals is from the surface of the sea...Health and Energetics of Polar Bears
Research in this focal area is centered on (i) collecting data on a variety of systems that help determine and mediate polar bear health and energetics, and (ii) developing monitoring and surveillance programs for detecting changes in population health over time. Additionally, this work will allow us to develop an understanding of how polar bear populations will respond to a variety of stressors... - Data
Metabolic Rate, Body Composition, and Blood Biochemistry Data from Polar Bears (Ursus maritimus) on Land, Western Hudson Bay, Canada, 2019-2022
This dataset is one table with time-linked behavior data derived from video camera collars on polar bears on land near Churchill, Manitoba, Canada. Eighteen polar bears were equipped with video-camera collars (Vertex Plus collar with camera option, Vectronic Aerospace GmbH, Berlin, Germany) for 19 - 23 days in 2019, 2021, and 2022.Polar Bear Continuous Time-Correlated Random Walk (CTCRW) Location Data Derived from Satellite Location Data, Chukchi and Beaufort Seas, July-November 1985-2017
his dataset consists of one table with estimated locations of adult female polar bears during July-November 1985-2017, used for quantifying changes in summer land use over time. Locations were estimated with a Continuous Time-Correlated Random Walk (CTCRW) model fit to satellite tracking from radio-collared adult female polar bears. All bears included in this data set were captured and instrumentePolar Bear Continuous Time-Correlated Random Walk (CTCRW) Location Data Derived from Satellite Location Data, Southern Beaufort Sea, 1986-2016
This dataset consists of one table with predicted locations of adult female polar bears. Locations were derived by a Continuous Time-Correlated Random Walk (CTCRW) model using satellite tracking radio-collared adult female polar bears captured and instrumented in the southern Beaufort Sea, 1986–2016.Serum Urea and Creatinine Levels of Spring-Caught Polar Bears (Ursus maritimus) in the Southern Beaufort and Chukchi Seas
These data are serum urea nitrogen and creatinine levels for polar bears captured in the southern Beaufort Sea 1983-2016 and the Chukchi Sea 1987-1993 and 2008-2017. The dataset includes relevant information about the bears that were captured including the latitude and longitude of their capture location, capture date, age class and sex, the age and number of cubs accompanying an adult female, andSatellite Location and Tri-axial Accelerometer Data from Adult Female Polar Bears (Ursus maritimus) in the Southern Beaufort Sea, April-October 2014
These data are from 5 adult female polar bears instrumented in the southern Beaufort Sea, April to October 2014. The dataset is comprised of two data packages: 1) contains GPS and Argos locations collected by satellite-linked GPS receivers mounted on external collars, and 2) contains archival logger data including measures of tri-axial acceleration and conductivity. These data were collected to gaLocations Collected 1985-2015 from Female Polar Bears (Ursus maritimus) with Dependent Young Instrumented in the Southern Beaufort Sea with Satellite-linked Transmitters by the USGS
This dataset contains a select subset of Argos and GPS locations collected by satellite data collection systems from collared adult female polar bears that were instrumented in the southern Beaufort Sea between 1985-2015. These data were collected to gain insights into movements of southern Beaufort Sea polar bears. These data were collected from adult female polar bears who had dependent young atBioelectrical Impedance, Deuterium Dilution, Body Mass, and Morphological Measures of Southern Beaufort Sea Female Polar Bears, Spring 2014-2016
This dataset contains data from the use of bioelectrical impedance analysis and deuterium injection as methods to estimate the body composition of female polar bears in the southern Beaufort Sea subpopulation. Data are provided on bioelectrical impedance resistance measures, the enrichment level of deuterium oxide that was injected and measured in blood samples, and morphological measures.Metabolic Rate, Body Composition, Foraging Success, Behavior, and GPS Locations of Female Polar Bears (Ursus maritimus), Beaufort Sea, Spring, 2014-2016 and Resting Energetics of an Adult Female Polar Bear
This data release comprises 4 datasets used to measure the field metabolic rate, body composition, foraging success, behavior, and movement patterns of 9 female polar bears on the sea ice of the Beaufort Sea in April, 2014-2016 as well as 1 dataset used to measure the energetic cost of resting in an adult female polar bear at the San Diego Zoo, San Diego, CA. Wild bears were dosed with and had theEnergetic Costs of Locomotion in Bears
Oxygen consumption and kinematic measurement data and tri-axial acceleration measurement data from 2 captive polar bears and 7 captive grizzly bears resting and walkingMeasures of oxygen consumption and stroke frequency of a captive subadult polar bear (Ursus maritimus) while resting in water and swimming and diving in a metabolic water flume, Oregon Zoo, 2017
This dataset contains measures of oxygen consumption and stroke frequency from 1 captive subadult female polar bear (166.5 kg) resting in the water (n = 7 sessions) and swimming and diving in a metabolic swim flume with water circulated at approximately 0.6 km/hr during swimming and diving measurements (n = 6 sessions) in September 2017. - Multimedia
POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022
This is B-roll video of POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022.
This is B-roll video of POV Polar Bear Collar Cam B-roll 2019, 2021, and 2022.
Polar bear with a GPS-enabled video camera collarAn adult male polar bear with a GPS-enabled video camera collar on land in Wapusk National Park, Canada as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
An adult male polar bear with a GPS-enabled video camera collar on land in Wapusk National Park, Canada as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Adult male polar bear interacting with two other bears in the oceanAdult male polar bear interacting with two other bears in the oceanPoint-of-view image of an adult male polar bear interacting with two other bears while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Adult male polar bear interacting with another bear in the oceanAdult male polar bear interacting with another bear in the oceanPoint-of-view image of an adult male polar bear interacting with another bear while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with another bear while in the ocean near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Adult male polar bear walking along the coastPoint-of-view image of an adult male polar bear walking along the coast in Wapusk National Park, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear walking along the coast in Wapusk National Park, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Adult male polar bear interacting with two other bears while on landAdult male polar bear interacting with two other bears while on landPoint-of-view image of an adult male polar bear interacting with two other bears while on land near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
Point-of-view image of an adult male polar bear interacting with two other bears while on land near Churchill, Manitoba, Canada. The image was recorded from a GPS-enabled video camera collar as part of a study measuring the energy expenditure, behavior, movement, and body composition changes of polar bears on land.
San Diego Zoo Polar BearTatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
San Diego Zoo Polar BearTatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
Tatqiq, an adult female polar bear at the San Diego Zoo, wearing an accelerometer collar being used to develop a method to remotely identify wild polar bear behaviors.
- Publications
Filter Total Items: 26
Ursids evolved dietary diversity without major alterations in metabolic rates
The diets of the eight species of ursids range from carnivory (e.g., polar bears, Ursus maritimus) to insectivory (e.g., sloth bears, Melursus ursinus), omnivory (e.g., brown bears, U. arctos), and herbivory (e.g., giant pandas, Ailuropoda melanoleuca). Dietary energy availability ranges from the high-fat, highly digestible, calorically dense diet of polar bears (~ 6.4 kcal digestible energy/g freAuthorsAnthony M. Carnahan, Anthony M. Pagano, Amelia L. Christian, Karyn D. Rode, Charles T. RobbinsPolar bear energetic and behavioral strategies on land with implications for surviving the ice-free period
Declining Arctic sea ice is increasing polar bear land use. Polar bears on land are thought to minimize activity to conserve energy. Here, we measure the daily energy expenditure (DEE), diet, behavior, movement, and body composition changes of 20 different polar bears on land over 19–23 days from August to September (2019–2022) in Manitoba, Canada. Polar bears on land exhibited a 5.2-fold range in
AuthorsAnthony M. Pagano, Karyn D. Rode, Nicholas J. Lunn, David McGeachy, Stephen N. Atkinson, Sean D. Farley, Joy A. Erlenbach, Charles T. RobbinsIncremental evolution of modeling a prognosis for polar bears in a rapidly changing Arctic
Updating predictions of the response of high-profile, at-risk species to climate change and anthropogenic stressors is vital for informing effective conservation action. Here, we review two prior generations of Bayesian network probability models predicting changes in global polar bear (Ursus maritimus) population status, and provide a contemporary update based on recent research findings and sea-AuthorsBruce G. Marcot, Todd C. Atwood, David C. Douglas, Jeffrey F. Bromaghin, Anthony M. Pagano, Steven C. AmstrupA body composition model with multiple storage compartments for polar bears (Ursus maritimus)
Climate warming is rapidly altering Arctic ecosystems. Polar bears (Ursus maritimus) need sea ice as a platform from which to hunt seals, but increased sea-ice loss is lengthening periods when bears are without access to primary hunting habitat. During periods of food scarcity, survival depends on the energy that a bear has stored in body reserves, termed storage energy, making this a key metric iAuthorsStephanie R. Penk, Pranav Sadana, Louise C. Archer, Anthony M. Pagano, Marc R. L. Cattet, Nicholas J. Lunn, Gregory W. Thiemann, Péter K. MolnárObserved and forecasted changes in land use by polar bears in the Beaufort and Chukchi Seas, 1985–2040
Monitoring changes in the distribution of large carnivores is important for managing human safety and supporting conservation. Throughout much of their range, polar bears (Ursus maritimus) are increasingly using terrestrial habitats in response to Arctic sea ice decline. Their increased presence in coastal areas has implications for bear-human conflict, inter-species interactions, and polar bear hAuthorsKaryn D. Rode, David C. Douglas, Todd C. Atwood, George M. Durner, Ryan R. Wilson, Anthony M. PaganoEffects of sea ice decline and summer land use on polar bear home range size in the Beaufort Sea
Animals responding to habitat loss and fragmentation may increase their home ranges to offset declines in localized resources or they may decrease their home ranges and switch to alternative resources. In many regions of the Arctic, polar bears (Ursus maritimus) exhibit some of the largest home ranges of any quadrupedal mammal. Polar bears are presently experiencing a rapid decline in Arctic sea iAuthorsAnthony M. Pagano, George M. Durner, Todd C. Atwood, David C. DouglasHuman-polar bear interactions
Human-wildlife interactions (HWI) are driven fundamentally by overlapping space and resources. As competition intensifies, the likelihood of interaction and conflict increases. In turn, conflict may impede conservation efforts by lowering social tolerance of wildlife, especially when human-wildlife conflict (HWC) poses a threat to human safety and economic well-being. Thus, mitigating conflict isAuthorsTodd C. Atwood, James WilderPolar bear foraging behavior
Polar bears forage in the marine environment, primarily on the sea ice over the shallow waters of the continental shelf. They are solitary, ambush hunters that catch ringed and bearded seals when they surface to breathe in ice holes or haul out on the ice to rest and molt. In most parts of their range, polar bears experience dramatic seasonal variability in their ability to catch seals, with foragAuthorsAnthony M. PaganoIdentifying reliable indicators of fitness in polar bears
Animal structural body size and condition are often measured to evaluate individual health, identify responses to environmental change and food availability, and relate food availability to effects on reproduction and survival. A variety of condition metrics have been developed but relationships between these metrics and vital rates are rarely validated. Identifying an optimal approach to estimateAuthorsKaryn D. Rode, Todd C. Atwood, Gregory Thiemann, Michelle St. Martin, Ryan H. Wilson, George M. Durner, Eric V. Regehr, Sandra L. Talbot, Kevin Sage, Anthony M. Pagano, Kristin S. SimacThe seasonal energetic landscape of an apex marine carnivore, the polar bear
Divergent movement strategies have enabled wildlife populations to adapt to environmental change. In recent decades, the Southern Beaufort Sea subpopulation of polar bears (Ursus maritimus) has developed a divergent movement strategy in response to diminishing sea ice where the majority of the subpopulation (73–85%) stays on the sea ice in summer and the remaining bears move to land. Although declAuthorsAnthony M. Pagano, Todd C. Atwood, George M. Durner, Terrie M. WilliamsOverhauling ocean spatial planning to improve marine megafauna conservation
Tracking data have led to evidence-based conservation of marine megafauna, but a disconnect remains between the many thousands of individual animals that have been tracked and the use of these data in conservation and management actions. Furthermore, the focus of most conservation efforts is within Exclusive Economic Zones despite the ability of these species to move thousands of kilometres acrossAuthorsAna M. M. Sequeira, Graeme C. Hays, David W. Sims, Victor M. Eguíluz, Jorge P. Rodríguez, Michelle R. Heupel, Robert G. Harcourt, Hannah J. Calich, Nuno Queiroz, Daniel P. Costa, Juan Fernández-Gracia, Luciana C. Ferreira, Simon D. Goldsworthy, Mark Hindell, Mary-Anne Lea, Mark G. Meekan, Anthony M. Pagano, Scott A. Shaffer, Julia Reisser, Michele Thums, Michael J Weise, Carlos M. DuarteEnergetic costs of aquatic locomotion in a subadult polar bear
Most marine mammals rely on swimming as their primary form of locomotion. These animals have evolved specialized morphologies, physiologies, and behaviors that have enabled them to efficiently move through an aquatic environment (Williams 1999). Such adaptations include body streamlining, modified plantar surfaces for propulsion, and abilities to remain submerged for extended durations (Williams 1AuthorsAnthony M. Pagano, Amy Cutting, Nicole Nicassio-Hiskey, Amy Hash, Terrie M. WilliamsNon-USGS Publications**
Pagano, A.M., A.M. Carnahan, C.T. Robbins, M.A. Owen, T. Batson, N. Wagner, A.
Cutting, N. Nicassio-Hiskey, A. Hash, and T.M. Williams. 2018. Energetic costs of
locomotion in bears: Is plantigrade locomotion energetically economical? Journal of
Experimental Biology vol. 221 no. 12 p.1-9. doi: 10.1242/jeb.175372Pagano, A.M., G.M. Durner, K.D. Rode, T.C. Atwood, S.N. Atkinson, E. Peacock, D.P.
Costa, M.A. Owen, and T.M. Williams. 2018. High-energy, high-fat lifestyle challenges an
Arctic apex predator, the polar bear. Science vol. 359 no. 6375 p. 568-572. doi: 10.1126/science.aan8677Pagano, A. M., Rode, K. D., Cutting, A., Owen, M. A., Jensen, S., Ware, J. V., Robbins, C. M., Durner, G. M., Atwood, T. C., Obbard, M. E., Middel, K. R., Thiemann, G. W., and Williams, T. M., 2017, Using tri-axial accelerometers to identify wild polar bear behaviors: Endangered Species Research vol. 32 p. 19-33. doi: 10.3354/esr00779Ware, J. V., Rode, K. D., Bromaghin, J. F., Douglas, D. C. Wilson, R. R., Regehr, E. V., Amstrup, S. C., Durner, G. M., Pagano, A. M., Olson, J. W., Robbins, C. T. and Jansen, H. T., 2017, Habitat degradation affects the summer activity of polar bears: Oecologia vol. 184 no. 1 p. 87-89. doi: 10.1007/s00442-017-3839-yWiig, Ø., Born, E. W., Laidre, K. L., Dietz, R., Jensen, M. V., Durner, G. M., Pagano, A. M., Regehr, E. V., St. Martin, M., Atkinson, S. N., and Dyck, M., 2017, Performance and retention of lightweight satellite radio tags applied to the ears of polar bears (Ursus maritimus): Animal Biotelemetry vol. 5 no. 9. doi: 10.1186/s40317-017-0124-0Sequeira, A.M.M., J.P. Rodríguez, V.M. Eguíluz, R. Harcourt, M. Hindell, D.W. Sims, C.M. Duarte, D.P. Costa, J. Fernández-Gracia, L.C. Ferreira, G.C. Hays, M.R. Heupel, M.G. Meekan, A. Aven, F. Bailleul, A.M.M. Baylis, M.L. Berumen, C.D. Braun, J. Burns, M.J. Caley, R. Campbell, R.H. Carmichael, E. Clua, L.D. Einoder, A. Friedlaender, M.E. Goebel, S.D. Goldsworthy, C. Guinet, J. Gunn, D. Hamer, N. Hammerschlag, M. Hammill, L.A. Hückstädt, N.E. Humphries, M.-A. Lea, A. Lowther, A. Mackay, E. McHuron, J. McKenzie, L. McLeay, C.R. McMahon, K. Mengersen, M.M.C. Muelbert, A. M. Pagano, B. Page, N. Queiroz, P.W. Robinson, S.A. Shaffer, M. Shivji, G.B. Skomal, S.R. Thorrold, S. Villegas-Amtmann, M. Weise, R. Wells, B. Wetherbee, A. Wiebkin, B. Wienecke, and M. Thums. 2018. Convergence of marine megafauna movement patterns in coastal and open oceans. Proceedings of the National Academy of Sciences vol. 115 no. 12 p.3072-3077. doi: 10.1073/pnas.1716137115Pagano, A.M., K.D. Rode, and S.N. Atkinson. 2017. Evaluating methods to assess the body condition of female polar bears. Ursus vol. 28 no. 2 p.171-181. doi: 10.2192/URSU-D-16-00029.1Pagano, A. M. and Arnold, T. W., 2009, Detection probabilities for ground-based breeding waterfowl surveys: Journal of Wildlife Management vol. 73 no. 3 p. 392-398. doi: 10.2193/2007-411Pagano, A. M. and Arnold, T. W., 2009, Estimating detection probabilities of waterfowl broods from ground-based surveys: Journal of Wildlife Management vol. 73 no. 5 p. 686-694. doi: 10.2193/2007-524Arnold, T. W., Pagano, A. M., Devries, J. H., Emery, R. B., Howerter, D. W., and Joynt, B. L., 2008, Social indices of breeding productivity in parkland mallards: Journal of Wildlife Management vol. 72 no. 1 p. 224-230. doi: 10.2193/2007-035**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
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