Page Klug


The common thread in my research has been the influence of human-induced,ecosystem modifications on species interactions. The human modifications I have investigated range from row-crop agriculture and rangeland management to the impact of introduced species. The majority of my research experience has focused on interactions between birds and their predators in human-altered landscapes. I have assessed how predators respond to landscape composition surrounding fragmented grasslands in an agricultural ecosystem and the impact on nesting birds. I have further investigated interactions between birds and snakes in a relatively contiguous tallgrass prairie by modeling the overlap of predator and prey habitat to illustrate how grazing and burning may exacerbate interactions. My current research investigates the effectiveness of control technologies to minimize the impact of brown treesnakes (Boiga irregularis) on the endangered Mariana Swiftlet (Aerodramus bartschi) on Guam. I perform additional research on the impact of invasive species on native fauna in southern Florida where invasive constrictor snakes and tegu lizards threaten local biodiversity. 



Ph.D. Biology, Kansas State University (2009)

M.S. Biology, University of Nebraska (2005)

B.S. Environmental Science and Policy, Drake University (2001)



Research Ecologist, US Geological Survey, Fort Collins Science Center (2013-present)

Assistant Director, University of Notre Dame Environmental Research Center - West (2010-2012) 

Graduate Teaching Assistant, Division of Biology, Kansas State University (2005-2009) 

Graduate Teaching/Research Assistant, Department of Biology, University of Nebraska (2003-2005)

Biological Project Assistant, Cary Institute of Ecosystem Studies (2002)

Biological Field Technician, University of Pittsburgh (2001)



American Society of Ichthyologists and Herpetologists

The National Chapter of The Wildlife Society

Cooper Ornithological Society








Janousek, W, PE Klug, JP McCarty, WA Cox, LI Berkeley, LL Wolfenbarger. 2015. The Conservation Value of Marginal Grasslands in a High-Intensity Agricultural Landscape for an Obligate Grassland Bird, the Dickcissel (Spiza americana). Journal of Wildlife Management. (In review).

McEachern, MA, AA Yackel Adams, PE Klug, LA Fitzgerald, RN Reed. 2015. Brumation of introduced Black and White Tegus, Tupinambis merianae (Squamata: Teiidae) in southern Florida. Southeastern Naturalist (In press).

Holldorf, ET, SR Siers, JQ Richmond, PE Klug, RN Reed. 2015. Invaded invaders: recent infection of invasive Brown Treesnakes on Guam by an exotic parasitic flatworm with a life cycle comprised of non-native hosts. PLoS ONE (In review).

Klug, PE, RN Reed, FJ Mazzotti, MA McEachern, JJ Vinci, KK Craven, AA Yackel Adams. 2015. The influence of disturbed habitat on the spatial ecology of Argentine tegu (Tupinambis merianae), a recent invader in the Everglades ecosystem (Florida, U.S.A.). Biological Invasions (DOI 10.1007/s10530-014-0834-7). [Download File]

Cox, WA, LL Wolfenbarger, LI Berkeley, SE Engberg, W Janousek, PE Klug, N Rider, JP McCarty. 2014. Grassland bird communities on conservation and marginal grasslands in an agricultural landscape. Agriculture, Ecosystems & Environment 193:53-59. [Download File]

Klug, PE, KA With, and SM. Wisley. 2011. Population genetic structure and landscape connectivity of the Eastern Yellowbelly Racer (Coluber constrictor flaviventris) in the contiguous tallgrass prairie of northeastern Kansas, USA. Landscape Ecology 26:281–294. [Download File]

Klug, PE, J Fill, and KA With. 2011. Spatial ecology of Eastern Yellowbelly Racer (Coluber constrictor flaviventris) and Great Plains Ratsnake (Pantherophis emoryi) in a contiguous tallgrass prairie landscape. Herpetologica 67(4): 428-439. [Download File]

Klug, PE, SL Jackrel, KA With. 2010. Linking snake habitat use to nest predation risk in grassland birds: the dangers of shrub cover. Oecologia 162: 803-813. [Download File]

Klug, PE, LL Wolfenbarger, and JP McCarty. 2010. Snakes are important nest predators of Dickcissels in an agricultural landscape. Wilson Journal of Ornithology 122(4): 799-803. [Download File]

Klug, PE, LL Wolfenbarger, and JP McCarty. 2009. The nest predator community of grassland birds responds to agroecosystem habitat at multiple scales. Ecography 32: 973-982. [Download File]