Study area of benthic macroinvertebrate response to Nisqually River Delta marsh restoration

Sampling sites are color-coded by habitat. White labels indicate site names. Marsh sites: U1, Unit 1; U2, Unit 2; U3, Unit 3; U4, Unit 4; M, Madrone; P1, Phase 1 restoration; P2, Phase 2 restoration; RF, reference. For Marsh sites, the last letter indicates the position along a tidal channel: N, north; M, middle; S, south; W, west. Tidal Flats site names all start with F. Elevation was determined from topobathymetric lidar data collected in 2011. The dashed line shows where dikes were removed. The star shows the location of the Billy Frank Junior National Wildlife Refuge headquarters.