Todd Atwood, Ph.D.
Todd is a Research Wildlife Biologist for the Alaska Science Center.
Professional Experience
2012 - Present Research Wildlife Biologist and Project Leader, USGS Alaska Science Center, Anchorage, AK
2008 - 2012 Research Wildlife Biologist, USDA/APHIS/WS/National Wildlife Research Center, Fort Collins, CO
2006 - 2008 Research Biologist, Research Branch, Arizona Game and Fish Department, Phoenix, AZ
2006 Biological Technician, USDA/National Wildlife Research Center, Fort Collins, CO
Education and Certifications
Ph.D. 2006 Utah State University Wildlife Biology
M.S. 2002 Purdue University Wildlife Ecology
B.S. 1999 Purdue University Wildlife Ecology
Affiliations and Memberships*
The Wildlife Society
International Association for Bear Research and Management
American Society of Mammalogists
Science and Products
Filter Total Items: 27
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Filter Total Items: 88
Measuring adrenal and reproductive hormones in hair from southern Beaufort Sea polar bears (Ursus maritimus) Measuring adrenal and reproductive hormones in hair from southern Beaufort Sea polar bears (Ursus maritimus)
Polar bears (Ursus maritimus) use sea ice to access marine mammal prey. In Alaska’s Southern Beaufort Sea, the declining availability of sea ice habitat in summer and fall has reduced opportunities for polar bears to routinely hunt on the ice for seals, their primary prey. This reduced access to prey may result in physiological stress with subsequent potential consequences to...
Authors
Marilize Van der Walt, Lorin Neuman-Lee, Patricia Terletzky, Todd Atwood, Eric Gese, Susannah French
How Is climate change affecting polar bears and giant pandas? How Is climate change affecting polar bears and giant pandas?
Anthropogenic greenhouse gas emissions are the primary cause of climate change and an estimated increase of 3.7 to 4.8 °C is predicted by the year 2100 if emissions continue at current levels. Polar bears (Ursus maritimus) and giant pandas (Ailuropoda melanoleuca) provide an interesting comparison study of the impact of climate change on bear species. While polar bears and giant pandas...
Authors
Melissa Songer, Todd Atwood, David Douglas, Qiongyu Huang, Renqiang Li, Nicholas Pilfold, Ming Xu, George Durner
Analyses on subpopulation abundance and annual number of maternal dens for the U.S. Fish and Wildlife Service on polar bears (Ursus maritimus) in the southern Beaufort Sea, Alaska Analyses on subpopulation abundance and annual number of maternal dens for the U.S. Fish and Wildlife Service on polar bears (Ursus maritimus) in the southern Beaufort Sea, Alaska
The long-term persistence of polar bears (Ursus maritimus) is threatened by sea-ice loss due to climate change, which is concurrently providing an opportunity in the Arctic for increased anthropogenic activities including natural resource extraction. Mitigating the risk of those activities, which can adversely affect the population dynamics of the southern Beaufort Sea (SBS)...
Authors
Todd Atwood, Jeffrey Bromaghin, Vijay Patil, George Durner, David Douglas, Kristin Simac
Identifying reliable indicators of fitness in polar bears Identifying reliable indicators of fitness in polar bears
Animal structural body size and condition are often measured to evaluate individual health, identify responses to environmental change and food availability, and relate food availability to effects on reproduction and survival. A variety of condition metrics have been developed but relationships between these metrics and vital rates are rarely validated. Identifying an optimal approach...
Authors
Karyn Rode, Todd Atwood, Gregory Thiemann, Michelle St. Martin, Ryan Wilson, George Durner, Eric Regehr, Sandra Talbot, Kevin Sage, Anthony Pagano, Kristin Simac
Fatty acid-based diet estimates suggest ringed seal remain the main prey of southern Beaufort Sea polar bears despite recent use of onshore food resources Fatty acid-based diet estimates suggest ringed seal remain the main prey of southern Beaufort Sea polar bears despite recent use of onshore food resources
Polar bears (Ursus maritimus) from the southern Beaufort Sea (SB) subpopulation have traditionally fed predominantly upon ice‐seals; however, as the proportion of the subpopulation using onshore habitat has recently increased, foraging on land‐based resources, including remains of subsistence‐harvested bowhead whales (Balaena mysticetus) and colonial nesting seabirds has been observed...
Authors
Jennifer Bourque, Todd Atwood, George Divoky, Connie Stewart, Melissa McKinney
Catalogue of polar bear (Ursus maritimus) maternal den locations in the Beaufort and Chukchi Seas and nearby areas, 1910–2018 Catalogue of polar bear (Ursus maritimus) maternal den locations in the Beaufort and Chukchi Seas and nearby areas, 1910–2018
This report presents data on the approximate locations and methods of discovery of 530 polar bear (Ursus maritimus) maternal dens observed in the Beaufort and Chukchi Seas and neighboring areas from 1910 to 2018, and archived partly by the U.S. Geological Survey, Alaska Science Center, and partly by the U.S. Fish and Wildlife Service, Marine Mammals Management, in Anchorage, Alaska. A...
Authors
George Durner, Steven Amstrup, Todd Atwood, David C. Douglas, Anthony S. Fischbach, Jay Olson, Karyn Rode, Ryan Wilson
The seasonal energetic landscape of an apex marine carnivore, the polar bear The seasonal energetic landscape of an apex marine carnivore, the polar bear
Divergent movement strategies have enabled wildlife populations to adapt to environmental change. In recent decades, the Southern Beaufort Sea subpopulation of polar bears (Ursus maritimus) has developed a divergent movement strategy in response to diminishing sea ice where the majority of the subpopulation (73–85%) stays on the sea ice in summer and the remaining bears move to land...
Authors
Anthony Pagano, Todd Atwood, George Durner, Terrie Williams
Use of subsistence-harvested whale carcasses by polar bears in the southern Beaufort Sea Use of subsistence-harvested whale carcasses by polar bears in the southern Beaufort Sea
The availability of a food subsidy has the potential to influence the condition, behavior, fitness, and population dynamics of a species. Since the early 2000s, monitoring efforts along the coast of northern Alaska indicated a higher proportion of polar bears (Ursus maritimus) of the southern Beaufort Sea (SB) subpopulation come onshore and feed at subsistence-harvested bowhead whale...
Authors
Kate Lillie, Eric Gese, Todd Atwood, Mary Conner
Reference intervals for blood-based biochemical analytes of southern Beaufort Sea polar bears Reference intervals for blood-based biochemical analytes of southern Beaufort Sea polar bears
Accurate reference intervals (RI) for commonly measured blood-based analytes are essential for health monitoring programs. Baseline values for a panel of analytes can be used to monitor physiologic and pathophysiologic processes such as organ function, electrolyte balance, and protein catabolism. Our reference population includes 651 serum samples from polar bears (Ursus maritimus) from...
Authors
Tricia Fry, Kristen Friedrichs, Todd Atwood, Colleen Duncan, Kristin Simac, Tony Goldberg
Global change-driven use of onshore habitat impacts polar bear faecal microbiota Global change-driven use of onshore habitat impacts polar bear faecal microbiota
The gut microbiota plays a critical role in host health, yet remains poorly studied in wild species. Polar bears (Ursus maritimus), key indicators of Arctic ecosystem health and environmental change, are currently affected by rapid shifts in habitat that may alter gut homeostasis. Declining sea ice has led to a divide in the southern Beaufort Sea polar bear subpopulation such that an...
Authors
Sophie Watson, Heidi Hauffe, Matthew Bull, Todd Atwood, Melissa McKinney, Massimo Pindo, Sarah Perkins
Are polar bear habitat resource selection functions developed from 1985-1996 data still useful? Are polar bear habitat resource selection functions developed from 1985-1996 data still useful?
1. Greenhouse gas-induced warming in the Arctic has caused declines in sea ice extent and changed its composition, raising concerns by all circumpolar nations for polar bear conservation. 2. Negative impacts have been observed in three well-studied polar bear subpopulations. Most subpopulations, however, receive little or no direct monitoring, hence, resource selection functions (RSF)...
Authors
George Durner, David C. Douglas, Todd Atwood
State of knowledge on current exposure, fate and potential health effects of contaminants in polar bears from the circumpolar Arctic State of knowledge on current exposure, fate and potential health effects of contaminants in polar bears from the circumpolar Arctic
The polar bear (Ursus maritimus) is among the Arctic species exposed to the highest concentrations of long-range transported bioaccumulative contaminants, such as halogenated organic compounds and mercury. Contaminant exposure is considered to be one of the largest threats to polar bears after the loss of their Arctic sea ice habitat due to climate change. The aim of this review is to...
Authors
Heli Routti, Todd Atwood, Thea Bechshoft, Andrei Boltunov, Tomasz Ciesielski, Jean-Pierre Desforges, Rune Dietz, Geir Gabrielsen, Bjorn Jenssen, Robert J. Letcher, Melissa McKinney, A. Morris, F. Riget, Christian Sonne, Bjarne Styrishave, Sabrina Tartu
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
Science and Products
Filter Total Items: 27
No Result Found
Filter Total Items: 88
Measuring adrenal and reproductive hormones in hair from southern Beaufort Sea polar bears (Ursus maritimus) Measuring adrenal and reproductive hormones in hair from southern Beaufort Sea polar bears (Ursus maritimus)
Polar bears (Ursus maritimus) use sea ice to access marine mammal prey. In Alaska’s Southern Beaufort Sea, the declining availability of sea ice habitat in summer and fall has reduced opportunities for polar bears to routinely hunt on the ice for seals, their primary prey. This reduced access to prey may result in physiological stress with subsequent potential consequences to...
Authors
Marilize Van der Walt, Lorin Neuman-Lee, Patricia Terletzky, Todd Atwood, Eric Gese, Susannah French
How Is climate change affecting polar bears and giant pandas? How Is climate change affecting polar bears and giant pandas?
Anthropogenic greenhouse gas emissions are the primary cause of climate change and an estimated increase of 3.7 to 4.8 °C is predicted by the year 2100 if emissions continue at current levels. Polar bears (Ursus maritimus) and giant pandas (Ailuropoda melanoleuca) provide an interesting comparison study of the impact of climate change on bear species. While polar bears and giant pandas...
Authors
Melissa Songer, Todd Atwood, David Douglas, Qiongyu Huang, Renqiang Li, Nicholas Pilfold, Ming Xu, George Durner
Analyses on subpopulation abundance and annual number of maternal dens for the U.S. Fish and Wildlife Service on polar bears (Ursus maritimus) in the southern Beaufort Sea, Alaska Analyses on subpopulation abundance and annual number of maternal dens for the U.S. Fish and Wildlife Service on polar bears (Ursus maritimus) in the southern Beaufort Sea, Alaska
The long-term persistence of polar bears (Ursus maritimus) is threatened by sea-ice loss due to climate change, which is concurrently providing an opportunity in the Arctic for increased anthropogenic activities including natural resource extraction. Mitigating the risk of those activities, which can adversely affect the population dynamics of the southern Beaufort Sea (SBS)...
Authors
Todd Atwood, Jeffrey Bromaghin, Vijay Patil, George Durner, David Douglas, Kristin Simac
Identifying reliable indicators of fitness in polar bears Identifying reliable indicators of fitness in polar bears
Animal structural body size and condition are often measured to evaluate individual health, identify responses to environmental change and food availability, and relate food availability to effects on reproduction and survival. A variety of condition metrics have been developed but relationships between these metrics and vital rates are rarely validated. Identifying an optimal approach...
Authors
Karyn Rode, Todd Atwood, Gregory Thiemann, Michelle St. Martin, Ryan Wilson, George Durner, Eric Regehr, Sandra Talbot, Kevin Sage, Anthony Pagano, Kristin Simac
Fatty acid-based diet estimates suggest ringed seal remain the main prey of southern Beaufort Sea polar bears despite recent use of onshore food resources Fatty acid-based diet estimates suggest ringed seal remain the main prey of southern Beaufort Sea polar bears despite recent use of onshore food resources
Polar bears (Ursus maritimus) from the southern Beaufort Sea (SB) subpopulation have traditionally fed predominantly upon ice‐seals; however, as the proportion of the subpopulation using onshore habitat has recently increased, foraging on land‐based resources, including remains of subsistence‐harvested bowhead whales (Balaena mysticetus) and colonial nesting seabirds has been observed...
Authors
Jennifer Bourque, Todd Atwood, George Divoky, Connie Stewart, Melissa McKinney
Catalogue of polar bear (Ursus maritimus) maternal den locations in the Beaufort and Chukchi Seas and nearby areas, 1910–2018 Catalogue of polar bear (Ursus maritimus) maternal den locations in the Beaufort and Chukchi Seas and nearby areas, 1910–2018
This report presents data on the approximate locations and methods of discovery of 530 polar bear (Ursus maritimus) maternal dens observed in the Beaufort and Chukchi Seas and neighboring areas from 1910 to 2018, and archived partly by the U.S. Geological Survey, Alaska Science Center, and partly by the U.S. Fish and Wildlife Service, Marine Mammals Management, in Anchorage, Alaska. A...
Authors
George Durner, Steven Amstrup, Todd Atwood, David C. Douglas, Anthony S. Fischbach, Jay Olson, Karyn Rode, Ryan Wilson
The seasonal energetic landscape of an apex marine carnivore, the polar bear The seasonal energetic landscape of an apex marine carnivore, the polar bear
Divergent movement strategies have enabled wildlife populations to adapt to environmental change. In recent decades, the Southern Beaufort Sea subpopulation of polar bears (Ursus maritimus) has developed a divergent movement strategy in response to diminishing sea ice where the majority of the subpopulation (73–85%) stays on the sea ice in summer and the remaining bears move to land...
Authors
Anthony Pagano, Todd Atwood, George Durner, Terrie Williams
Use of subsistence-harvested whale carcasses by polar bears in the southern Beaufort Sea Use of subsistence-harvested whale carcasses by polar bears in the southern Beaufort Sea
The availability of a food subsidy has the potential to influence the condition, behavior, fitness, and population dynamics of a species. Since the early 2000s, monitoring efforts along the coast of northern Alaska indicated a higher proportion of polar bears (Ursus maritimus) of the southern Beaufort Sea (SB) subpopulation come onshore and feed at subsistence-harvested bowhead whale...
Authors
Kate Lillie, Eric Gese, Todd Atwood, Mary Conner
Reference intervals for blood-based biochemical analytes of southern Beaufort Sea polar bears Reference intervals for blood-based biochemical analytes of southern Beaufort Sea polar bears
Accurate reference intervals (RI) for commonly measured blood-based analytes are essential for health monitoring programs. Baseline values for a panel of analytes can be used to monitor physiologic and pathophysiologic processes such as organ function, electrolyte balance, and protein catabolism. Our reference population includes 651 serum samples from polar bears (Ursus maritimus) from...
Authors
Tricia Fry, Kristen Friedrichs, Todd Atwood, Colleen Duncan, Kristin Simac, Tony Goldberg
Global change-driven use of onshore habitat impacts polar bear faecal microbiota Global change-driven use of onshore habitat impacts polar bear faecal microbiota
The gut microbiota plays a critical role in host health, yet remains poorly studied in wild species. Polar bears (Ursus maritimus), key indicators of Arctic ecosystem health and environmental change, are currently affected by rapid shifts in habitat that may alter gut homeostasis. Declining sea ice has led to a divide in the southern Beaufort Sea polar bear subpopulation such that an...
Authors
Sophie Watson, Heidi Hauffe, Matthew Bull, Todd Atwood, Melissa McKinney, Massimo Pindo, Sarah Perkins
Are polar bear habitat resource selection functions developed from 1985-1996 data still useful? Are polar bear habitat resource selection functions developed from 1985-1996 data still useful?
1. Greenhouse gas-induced warming in the Arctic has caused declines in sea ice extent and changed its composition, raising concerns by all circumpolar nations for polar bear conservation. 2. Negative impacts have been observed in three well-studied polar bear subpopulations. Most subpopulations, however, receive little or no direct monitoring, hence, resource selection functions (RSF)...
Authors
George Durner, David C. Douglas, Todd Atwood
State of knowledge on current exposure, fate and potential health effects of contaminants in polar bears from the circumpolar Arctic State of knowledge on current exposure, fate and potential health effects of contaminants in polar bears from the circumpolar Arctic
The polar bear (Ursus maritimus) is among the Arctic species exposed to the highest concentrations of long-range transported bioaccumulative contaminants, such as halogenated organic compounds and mercury. Contaminant exposure is considered to be one of the largest threats to polar bears after the loss of their Arctic sea ice habitat due to climate change. The aim of this review is to...
Authors
Heli Routti, Todd Atwood, Thea Bechshoft, Andrei Boltunov, Tomasz Ciesielski, Jean-Pierre Desforges, Rune Dietz, Geir Gabrielsen, Bjorn Jenssen, Robert J. Letcher, Melissa McKinney, A. Morris, F. Riget, Christian Sonne, Bjarne Styrishave, Sabrina Tartu
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
*Disclaimer: Listing outside positions with professional scientific organizations on this Staff Profile are for informational purposes only and do not constitute an endorsement of those professional scientific organizations or their activities by the USGS, Department of the Interior, or U.S. Government