Todd Atwood, Ph.D.
Todd is a Research Wildlife Biologist for the Alaska Science Center.
Professional Experience
2012 - Present Research Wildlife Biologist and Project Leader, USGS Alaska Science Center, Anchorage, AK
2008 - 2012 Research Wildlife Biologist, USDA/APHIS/WS/National Wildlife Research Center, Fort Collins, CO
2006 - 2008 Research Biologist, Research Branch, Arizona Game and Fish Department, Phoenix, AZ
2006 Biological Technician, USDA/National Wildlife Research Center, Fort Collins, CO
Education and Certifications
Ph.D. 2006 Utah State University Wildlife Biology
M.S. 2002 Purdue University Wildlife Ecology
B.S. 1999 Purdue University Wildlife Ecology
Affiliations and Memberships*
The Wildlife Society
International Association for Bear Research and Management
American Society of Mammalogists
Science and Products
Filter Total Items: 27
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Filter Total Items: 91
Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land
In response to a changing climate, many species alter habitat use. Polar bears Ursus maritimus in the southern Beaufort Sea have increasingly used land for maternal denning. To aid in detecting denning behavior, we developed an objective method to identify polar bear denning events using temperature sensor data collected by satellite-linked transmitters deployed on adult females between...
Authors
Jay W Olson, Karyn D. Rode, Dennis L. Eggett, T. S. Smith, R. R. Wilson, George M. Durner, Anthony S. Fischbach, Todd C. Atwood, David C. Douglas
Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines
Loss of Arctic sea ice owing to climate change is the primary threat to polar bears throughout their range. We evaluated the potential response of polar bears to sea-ice declines by (i) calculating generation length (GL) for the species, which determines the timeframe for conservation assessments; (ii) developing a standardized sea-ice metric representing important habitat; and (iii)...
Authors
Eric V. Regehr, Kristin L. Laidre, H. Resit Akcakaya, Steven C. Amstrup, Todd C. Atwood, Nicholas J. Lunn, Martyn E. Obbard, Harry Stern, Gregory W. Thiemann, Øystein Wiig
Forecasting the relative influence of environmental and anthropogenic stressors on polar bears Forecasting the relative influence of environmental and anthropogenic stressors on polar bears
Effective conservation planning requires understanding and ranking threats to wildlife populations. We developed a Bayesian network model to evaluate the relative influence of environmental and anthropogenic stressors, and their mitigation, on the persistence of polar bears (Ursus maritimus). Overall sea ice conditions, affected by rising global temperatures, were the most influential...
Authors
Todd C. Atwood, Bruce G. Marcot, David C. Douglas, Steven C. Amstrup, Karyn D. Rode, George M. Durner, Jeffrey F. Bromaghin
Rapid environmental change drives increased land use by an Arctic marine predator Rapid environmental change drives increased land use by an Arctic marine predator
In the Arctic Ocean’s southern Beaufort Sea (SB), the length of the sea ice melt season (i.e., period between the onset of sea ice break-up in summer and freeze-up in fall) has increased substantially since the late 1990s. Historically, polar bears (Ursus maritimus) of the SB have mostly remained on the sea ice year-round (except for those that came ashore to den), but recent changes in...
Authors
Todd C. Atwood, Elizabeth L. Peacock, Melissa A. McKinney, Kate Lillie, Ryan H. Wilson, David C. Douglas, Pat Terletzky, Susanne Miller
Testing methods for using high-resolution satellite imagery to monitor polar bear abundance and distribution Testing methods for using high-resolution satellite imagery to monitor polar bear abundance and distribution
High-resolution satellite imagery is a promising tool for providing coarse information about polar species abundance and distribution, but current applications are limited. With polar bears (Ursus maritimus), the technique has only proven effective on landscapes with little topographic relief that are devoid of snow and ice, and time-consuming manual review of imagery is required to...
Authors
Michelle A. LaRue, Seth P. Stapleton, Claire Porter, Stephen N. Atkinson, Todd C. Atwood, Markus Dyck, Nicolas Lecomte
Gene transcription in polar bears (Ursus maritimus) from disparate populations Gene transcription in polar bears (Ursus maritimus) from disparate populations
Polar bears in the Beaufort (SB) and Chukchi (CS) Seas experience different environments due primarily to a longer history of sea ice loss in the Beaufort Sea. Ecological differences have been identified as a possible reason for the generally poorer body condition and reproduction of Beaufort polar bears compared to those from the Chukchi, but the influence of exposure to other stressors...
Authors
Lizabeth Bowen, A. Keith Miles, Shannon C. Waters-Dynes, Randi Meyerson, Karyn D. Rode, Todd C. Atwood
Changing Arctic Ecosystems: Updated forecast: Reducing carbon dioxide (CO2) emissions required to improve polar bear outlook Changing Arctic Ecosystems: Updated forecast: Reducing carbon dioxide (CO2) emissions required to improve polar bear outlook
The Arctic is warming faster than other regions of the world due to the loss of snow and ice, which increases the amount of solar energy absorbed by the region. The most visible consequence has been the rapid decline in sea ice over the last 3 decades-a decline projected to bring long ice-free summers if greenhouse gas (GHG) emissions are not significantly reduced. The polar bear (Ursus...
Authors
Karen L. Oakley, Todd C. Atwood, Douglas N. Mugel, Karyn D. Rode, Mary E. Whalen
Enhanced biological processes associated with alopecia in polar bears (Ursus maritimus) Enhanced biological processes associated with alopecia in polar bears (Ursus maritimus)
Populations of wildlife species worldwide experience incidents of mass morbidity and mortality. Primary or secondary drivers of these events may escape classical detection methods for identifying microbial insults, toxin exposure, or additional stressors. In 2012, 28% of polar bears sampled in a study in the southern Beaufort Sea region of Alaska had varying degrees of alopecia that was
Authors
Lizabeth Bowen, A. Keith Miles, Jeffrey L. Stott, Shannon C. Waters-Dynes, Todd C. Atwood
Forecasting wildlife response to rapid warming in the Alaskan Arctic Forecasting wildlife response to rapid warming in the Alaskan Arctic
Arctic wildlife species face a dynamic and increasingly novel environment because of climate warming and the associated increase in human activity. Both marine and terrestrial environments are undergoing rapid environmental shifts, including loss of sea ice, permafrost degradation, and altered biogeochemical fluxes. Forecasting wildlife responses to climate change can facilitate...
Authors
Caroline R. Van Hemert, Paul L. Flint, Mark S. Udevitz, Joshua C. Koch, Todd C. Atwood, Karen L. Oakley, John M. Pearce
Polar bear population dynamics in the southern Beaufort Sea during a period of sea ice decline Polar bear population dynamics in the southern Beaufort Sea during a period of sea ice decline
In the southern Beaufort Sea of the United States and Canada, prior investigations have linked declines in summer sea ice to reduced physical condition, growth, and survival of polar bears (Ursus maritimus). Combined with projections of population decline due to continued climate warming and the ensuing loss of sea ice habitat, those findings contributed to the 2008 decision to list the...
Authors
Jeffrey F. Bromaghin, Trent L. McDonald, Ian Stirling, Andrew E. Derocher, Evan S. Richardson, Eric V. Regehr, David C. Douglas, George M. Durner, Todd C. Atwood, Steven C. Amstrup
A review of infectious agents in polar bears (Ursus maritimus) and their long-term ecological relevance A review of infectious agents in polar bears (Ursus maritimus) and their long-term ecological relevance
Disease was a listing criterion for the polar bear (Ursus maritimus) as threatened under the Endangered Species Act in 2008; it is therefore important to evaluate the current state of knowledge and identify any information gaps pertaining to diseases in polar bears. We conducted a systematic literature review focused on infectious agents and associated health impacts identified in polar...
Authors
Anna C. Fagre, Kelly A. Patyk, Pauline Nol, Todd C. Atwood, Karsten Hueffer, Colleen G. Duncan
Establishing a definition of polar bear (Ursus maritimus) health: A guide to research and management activities Establishing a definition of polar bear (Ursus maritimus) health: A guide to research and management activities
The meaning of health for wildlife and perspectives on how to assess and measure health, are not well characterized. For wildlife at risk, such as some polar bear (Ursus maritimus) subpopulations, establishing comprehensive monitoring programs that include health status is an emerging need. Environmental changes, especially loss of sea ice habitat, have raised concern about polar bear...
Authors
Kelly A. Patyk, Colleen G. Duncan, Pauline Nol, C. Sonne, Kristin L. Laidre, Martyn E. Obbard, Øystein Wiig, Jon Aars, Eric V. Regehr, L. Gustafson, Todd C. Atwood
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
Science and Products
Filter Total Items: 27
No Result Found
Filter Total Items: 91
Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land
In response to a changing climate, many species alter habitat use. Polar bears Ursus maritimus in the southern Beaufort Sea have increasingly used land for maternal denning. To aid in detecting denning behavior, we developed an objective method to identify polar bear denning events using temperature sensor data collected by satellite-linked transmitters deployed on adult females between...
Authors
Jay W Olson, Karyn D. Rode, Dennis L. Eggett, T. S. Smith, R. R. Wilson, George M. Durner, Anthony S. Fischbach, Todd C. Atwood, David C. Douglas
Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines
Loss of Arctic sea ice owing to climate change is the primary threat to polar bears throughout their range. We evaluated the potential response of polar bears to sea-ice declines by (i) calculating generation length (GL) for the species, which determines the timeframe for conservation assessments; (ii) developing a standardized sea-ice metric representing important habitat; and (iii)...
Authors
Eric V. Regehr, Kristin L. Laidre, H. Resit Akcakaya, Steven C. Amstrup, Todd C. Atwood, Nicholas J. Lunn, Martyn E. Obbard, Harry Stern, Gregory W. Thiemann, Øystein Wiig
Forecasting the relative influence of environmental and anthropogenic stressors on polar bears Forecasting the relative influence of environmental and anthropogenic stressors on polar bears
Effective conservation planning requires understanding and ranking threats to wildlife populations. We developed a Bayesian network model to evaluate the relative influence of environmental and anthropogenic stressors, and their mitigation, on the persistence of polar bears (Ursus maritimus). Overall sea ice conditions, affected by rising global temperatures, were the most influential...
Authors
Todd C. Atwood, Bruce G. Marcot, David C. Douglas, Steven C. Amstrup, Karyn D. Rode, George M. Durner, Jeffrey F. Bromaghin
Rapid environmental change drives increased land use by an Arctic marine predator Rapid environmental change drives increased land use by an Arctic marine predator
In the Arctic Ocean’s southern Beaufort Sea (SB), the length of the sea ice melt season (i.e., period between the onset of sea ice break-up in summer and freeze-up in fall) has increased substantially since the late 1990s. Historically, polar bears (Ursus maritimus) of the SB have mostly remained on the sea ice year-round (except for those that came ashore to den), but recent changes in...
Authors
Todd C. Atwood, Elizabeth L. Peacock, Melissa A. McKinney, Kate Lillie, Ryan H. Wilson, David C. Douglas, Pat Terletzky, Susanne Miller
Testing methods for using high-resolution satellite imagery to monitor polar bear abundance and distribution Testing methods for using high-resolution satellite imagery to monitor polar bear abundance and distribution
High-resolution satellite imagery is a promising tool for providing coarse information about polar species abundance and distribution, but current applications are limited. With polar bears (Ursus maritimus), the technique has only proven effective on landscapes with little topographic relief that are devoid of snow and ice, and time-consuming manual review of imagery is required to...
Authors
Michelle A. LaRue, Seth P. Stapleton, Claire Porter, Stephen N. Atkinson, Todd C. Atwood, Markus Dyck, Nicolas Lecomte
Gene transcription in polar bears (Ursus maritimus) from disparate populations Gene transcription in polar bears (Ursus maritimus) from disparate populations
Polar bears in the Beaufort (SB) and Chukchi (CS) Seas experience different environments due primarily to a longer history of sea ice loss in the Beaufort Sea. Ecological differences have been identified as a possible reason for the generally poorer body condition and reproduction of Beaufort polar bears compared to those from the Chukchi, but the influence of exposure to other stressors...
Authors
Lizabeth Bowen, A. Keith Miles, Shannon C. Waters-Dynes, Randi Meyerson, Karyn D. Rode, Todd C. Atwood
Changing Arctic Ecosystems: Updated forecast: Reducing carbon dioxide (CO2) emissions required to improve polar bear outlook Changing Arctic Ecosystems: Updated forecast: Reducing carbon dioxide (CO2) emissions required to improve polar bear outlook
The Arctic is warming faster than other regions of the world due to the loss of snow and ice, which increases the amount of solar energy absorbed by the region. The most visible consequence has been the rapid decline in sea ice over the last 3 decades-a decline projected to bring long ice-free summers if greenhouse gas (GHG) emissions are not significantly reduced. The polar bear (Ursus...
Authors
Karen L. Oakley, Todd C. Atwood, Douglas N. Mugel, Karyn D. Rode, Mary E. Whalen
Enhanced biological processes associated with alopecia in polar bears (Ursus maritimus) Enhanced biological processes associated with alopecia in polar bears (Ursus maritimus)
Populations of wildlife species worldwide experience incidents of mass morbidity and mortality. Primary or secondary drivers of these events may escape classical detection methods for identifying microbial insults, toxin exposure, or additional stressors. In 2012, 28% of polar bears sampled in a study in the southern Beaufort Sea region of Alaska had varying degrees of alopecia that was
Authors
Lizabeth Bowen, A. Keith Miles, Jeffrey L. Stott, Shannon C. Waters-Dynes, Todd C. Atwood
Forecasting wildlife response to rapid warming in the Alaskan Arctic Forecasting wildlife response to rapid warming in the Alaskan Arctic
Arctic wildlife species face a dynamic and increasingly novel environment because of climate warming and the associated increase in human activity. Both marine and terrestrial environments are undergoing rapid environmental shifts, including loss of sea ice, permafrost degradation, and altered biogeochemical fluxes. Forecasting wildlife responses to climate change can facilitate...
Authors
Caroline R. Van Hemert, Paul L. Flint, Mark S. Udevitz, Joshua C. Koch, Todd C. Atwood, Karen L. Oakley, John M. Pearce
Polar bear population dynamics in the southern Beaufort Sea during a period of sea ice decline Polar bear population dynamics in the southern Beaufort Sea during a period of sea ice decline
In the southern Beaufort Sea of the United States and Canada, prior investigations have linked declines in summer sea ice to reduced physical condition, growth, and survival of polar bears (Ursus maritimus). Combined with projections of population decline due to continued climate warming and the ensuing loss of sea ice habitat, those findings contributed to the 2008 decision to list the...
Authors
Jeffrey F. Bromaghin, Trent L. McDonald, Ian Stirling, Andrew E. Derocher, Evan S. Richardson, Eric V. Regehr, David C. Douglas, George M. Durner, Todd C. Atwood, Steven C. Amstrup
A review of infectious agents in polar bears (Ursus maritimus) and their long-term ecological relevance A review of infectious agents in polar bears (Ursus maritimus) and their long-term ecological relevance
Disease was a listing criterion for the polar bear (Ursus maritimus) as threatened under the Endangered Species Act in 2008; it is therefore important to evaluate the current state of knowledge and identify any information gaps pertaining to diseases in polar bears. We conducted a systematic literature review focused on infectious agents and associated health impacts identified in polar...
Authors
Anna C. Fagre, Kelly A. Patyk, Pauline Nol, Todd C. Atwood, Karsten Hueffer, Colleen G. Duncan
Establishing a definition of polar bear (Ursus maritimus) health: A guide to research and management activities Establishing a definition of polar bear (Ursus maritimus) health: A guide to research and management activities
The meaning of health for wildlife and perspectives on how to assess and measure health, are not well characterized. For wildlife at risk, such as some polar bear (Ursus maritimus) subpopulations, establishing comprehensive monitoring programs that include health status is an emerging need. Environmental changes, especially loss of sea ice habitat, have raised concern about polar bear...
Authors
Kelly A. Patyk, Colleen G. Duncan, Pauline Nol, C. Sonne, Kristin L. Laidre, Martyn E. Obbard, Øystein Wiig, Jon Aars, Eric V. Regehr, L. Gustafson, Todd C. Atwood
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
*Disclaimer: Listing outside positions with professional scientific organizations on this Staff Profile are for informational purposes only and do not constitute an endorsement of those professional scientific organizations or their activities by the USGS, Department of the Interior, or U.S. Government