Todd Atwood, Ph.D.
Todd is a Research Wildlife Biologist for the Alaska Science Center.
Professional Experience
2012 - Present Research Wildlife Biologist and Project Leader, USGS Alaska Science Center, Anchorage, AK
2008 - 2012 Research Wildlife Biologist, USDA/APHIS/WS/National Wildlife Research Center, Fort Collins, CO
2006 - 2008 Research Biologist, Research Branch, Arizona Game and Fish Department, Phoenix, AZ
2006 Biological Technician, USDA/National Wildlife Research Center, Fort Collins, CO
Education and Certifications
Ph.D. 2006 Utah State University Wildlife Biology
M.S. 2002 Purdue University Wildlife Ecology
B.S. 1999 Purdue University Wildlife Ecology
Affiliations and Memberships*
The Wildlife Society
International Association for Bear Research and Management
American Society of Mammalogists
Science and Products
Filter Total Items: 27
No Result Found
Filter Total Items: 88
Temporal complexity of southern Beaufort Sea polar bear diets during a period of increasing land use Temporal complexity of southern Beaufort Sea polar bear diets during a period of increasing land use
From 2000 to 2013, use of land as a seasonal habitat by polar bears (Ursus maritimus) of the southern Beaufort Sea (SB) subpopulation substantially increased. This onshore use has been linked to reduced spatial and temporal availability of sea ice, as well as to the availability of subsistence‐harvested bowhead whale (Balaena mysticetus) bone piles. Here, we evaluated the role of climate
Authors
Melissa McKinney, Todd Atwood, Sara Iverson, Elizabeth Peacock
Polar bear attacks on humans: Implications of a changing climate Polar bear attacks on humans: Implications of a changing climate
Understanding causes of polar bear (Ursus maritimus) attacks on humans is critical to ensuring both human safety and polar bear conservation. Although considerable attention has been focused on understanding black (U. americanus) and grizzly (U. arctos) bear conflicts with humans, there have been few attempts to systematically collect, analyze, and interpret available information on...
Authors
James Wilder, Dag Vongraven, Todd Atwood, Bob Hansen, Amalie Jessen, Anatoly Kochnev, Geoff York, Rachel Vallender, Daryll Hedman, Melissa Gibbons
Relative influences of climate change and human activity on the onshore distribution of polar bears Relative influences of climate change and human activity on the onshore distribution of polar bears
Climate change is altering habitat for many species, leading to shifts in distributions that can increase levels of human-wildlife conflict. To develop effective strategies for minimizing human-wildlife conflict, we must understand the relative influences that climate change and other factors have on wildlife distributions. Polar bears (Ursus maritimus) are increasingly using land during...
Authors
Ryan Wilson, Eric Regehr, Michelle St. Martin, Todd Atwood, Elizabeth Peacock, Susanne Miller, George Divoky
Demographic and temporal variations in immunity and condition of polar bears (Ursus maritimus) from the southern Beaufort Sea Demographic and temporal variations in immunity and condition of polar bears (Ursus maritimus) from the southern Beaufort Sea
Assessing the health and condition of animals in their natural environment can be problematic. Many physiological metrics, including immunity, are highly influenced by specific context and recent events to which researchers may be unaware. Thus, using a multifaceted physiological approach and a context-specific analysis encompassing multiple time scales can be highly informative...
Authors
Lorin Neuman-Lee, Patricia Terletzky, Todd Atwood, Eric Gese, Geoffrey Smith, Sydney Greenfield, John Pettit, Susannah French
Environmental and behavioral changes may influence the exposure of an Arctic apex predator to pathogens and contaminants Environmental and behavioral changes may influence the exposure of an Arctic apex predator to pathogens and contaminants
Recent decline of sea ice habitat has coincided with increased use of land by polar bears (Ursus maritimus) from the southern Beaufort Sea (SB), which may alter the risks of exposure to pathogens and contaminants. We assayed blood samples from SB polar bears to assess prior exposure to the pathogens Brucella spp., Toxoplasma gondii, Coxiella burnetii, Francisella tularensis, and Neospora...
Authors
Todd Atwood, Colleen Duncan, Kelly Patyk, Pauline Nol, Jack Rhyan, Matthew McCollum, Melissa McKinney, Andrew Ramey, Camila Cerqueira-Cezar, Oliver C H Kwok, Jitender Dubey, S.G. Hennager
Polar bears and sea ice habitat change Polar bears and sea ice habitat change
The polar bear (Ursus maritimus) is an obligate apex predator of Arctic sea ice and as such can be affected by climate warming-induced changes in the extent and composition of pack ice and its impacts on their seal prey. Sea ice declines have negatively impacted some polar bear subpopulations through reduced energy input because of loss of hunting habitats, higher energy costs due to...
Authors
George Durner, Todd Atwood
Uncertainties in forecasting the response of polar bears to global climate change Uncertainties in forecasting the response of polar bears to global climate change
Several sources of uncertainty affect how precisely the future status of polar bears (Ursus maritimus) can be forecasted. Foremost are unknowns about the future levels of global greenhouse gas emissions, which could range from an unabated increase to an aggressively mitigated reduction. Uncertainties also arise because different climate models project different amounts and rates of...
Authors
David Douglas, Todd Atwood
Ecological change drives a decline in mercury concentrations in southern Beaufort Sea polar bears Ecological change drives a decline in mercury concentrations in southern Beaufort Sea polar bears
We evaluated total mercury (THg) concentrations and trends in polar bears from the southern Beaufort Sea subpopulation from 2004 to 2011. Hair THg concentrations ranged widely among individuals from 0.6 to 13.3 μg g–1 dry weight (mean: 3.5 ± 0.2 μg g–1). Concentrations differed among sex and age classes: solitary adult females ≈ adult females with cubs ≈ subadults > adult males ≈...
Authors
Melissa McKinney, Todd Atwood, Sara Pedro, Elizabeth Peacock
Using tri-axial accelerometers to identify wild polar bear behaviors Using tri-axial accelerometers to identify wild polar bear behaviors
Tri-axial accelerometers have been used to remotely identify the behaviors of a wide range of taxa. Assigning behaviors to accelerometer data often involves the use of captive animals or surrogate species, as their accelerometer signatures are generally assumed to be similar to those of their wild counterparts. However, this has rarely been tested. Validated accelerometer data are needed...
Authors
Anthony Pagano, Karyn Rode, A. Cutting, M.A. Owen, S. Jensen, J.V. Ware, C.T. Robbins, George Durner, Todd Atwood, M.E. Obbard, K.R. Middel, G.W. Thiemann, T.M. Williams
Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land
In response to a changing climate, many species alter habitat use. Polar bears Ursus maritimus in the southern Beaufort Sea have increasingly used land for maternal denning. To aid in detecting denning behavior, we developed an objective method to identify polar bear denning events using temperature sensor data collected by satellite-linked transmitters deployed on adult females between...
Authors
Jay Olson, Karyn Rode, Dennis Eggett, T. Smith, R. Wilson, George Durner, Anthony S. Fischbach, Todd Atwood, David C. Douglas
Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines
Loss of Arctic sea ice owing to climate change is the primary threat to polar bears throughout their range. We evaluated the potential response of polar bears to sea-ice declines by (i) calculating generation length (GL) for the species, which determines the timeframe for conservation assessments; (ii) developing a standardized sea-ice metric representing important habitat; and (iii)...
Authors
Eric Regehr, Kristin Laidre, H. Akcakaya, Steven Amstrup, Todd Atwood, Nicholas J. Lunn, Martyn Obbard, Harry Stern, Gregory Thiemann, Øystein Wiig
Forecasting the relative influence of environmental and anthropogenic stressors on polar bears Forecasting the relative influence of environmental and anthropogenic stressors on polar bears
Effective conservation planning requires understanding and ranking threats to wildlife populations. We developed a Bayesian network model to evaluate the relative influence of environmental and anthropogenic stressors, and their mitigation, on the persistence of polar bears (Ursus maritimus). Overall sea ice conditions, affected by rising global temperatures, were the most influential...
Authors
Todd Atwood, Bruce Marcot, David Douglas, Steven Amstrup, Karyn Rode, George Durner, Jeffrey Bromaghin
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
Science and Products
Filter Total Items: 27
No Result Found
Filter Total Items: 88
Temporal complexity of southern Beaufort Sea polar bear diets during a period of increasing land use Temporal complexity of southern Beaufort Sea polar bear diets during a period of increasing land use
From 2000 to 2013, use of land as a seasonal habitat by polar bears (Ursus maritimus) of the southern Beaufort Sea (SB) subpopulation substantially increased. This onshore use has been linked to reduced spatial and temporal availability of sea ice, as well as to the availability of subsistence‐harvested bowhead whale (Balaena mysticetus) bone piles. Here, we evaluated the role of climate
Authors
Melissa McKinney, Todd Atwood, Sara Iverson, Elizabeth Peacock
Polar bear attacks on humans: Implications of a changing climate Polar bear attacks on humans: Implications of a changing climate
Understanding causes of polar bear (Ursus maritimus) attacks on humans is critical to ensuring both human safety and polar bear conservation. Although considerable attention has been focused on understanding black (U. americanus) and grizzly (U. arctos) bear conflicts with humans, there have been few attempts to systematically collect, analyze, and interpret available information on...
Authors
James Wilder, Dag Vongraven, Todd Atwood, Bob Hansen, Amalie Jessen, Anatoly Kochnev, Geoff York, Rachel Vallender, Daryll Hedman, Melissa Gibbons
Relative influences of climate change and human activity on the onshore distribution of polar bears Relative influences of climate change and human activity on the onshore distribution of polar bears
Climate change is altering habitat for many species, leading to shifts in distributions that can increase levels of human-wildlife conflict. To develop effective strategies for minimizing human-wildlife conflict, we must understand the relative influences that climate change and other factors have on wildlife distributions. Polar bears (Ursus maritimus) are increasingly using land during...
Authors
Ryan Wilson, Eric Regehr, Michelle St. Martin, Todd Atwood, Elizabeth Peacock, Susanne Miller, George Divoky
Demographic and temporal variations in immunity and condition of polar bears (Ursus maritimus) from the southern Beaufort Sea Demographic and temporal variations in immunity and condition of polar bears (Ursus maritimus) from the southern Beaufort Sea
Assessing the health and condition of animals in their natural environment can be problematic. Many physiological metrics, including immunity, are highly influenced by specific context and recent events to which researchers may be unaware. Thus, using a multifaceted physiological approach and a context-specific analysis encompassing multiple time scales can be highly informative...
Authors
Lorin Neuman-Lee, Patricia Terletzky, Todd Atwood, Eric Gese, Geoffrey Smith, Sydney Greenfield, John Pettit, Susannah French
Environmental and behavioral changes may influence the exposure of an Arctic apex predator to pathogens and contaminants Environmental and behavioral changes may influence the exposure of an Arctic apex predator to pathogens and contaminants
Recent decline of sea ice habitat has coincided with increased use of land by polar bears (Ursus maritimus) from the southern Beaufort Sea (SB), which may alter the risks of exposure to pathogens and contaminants. We assayed blood samples from SB polar bears to assess prior exposure to the pathogens Brucella spp., Toxoplasma gondii, Coxiella burnetii, Francisella tularensis, and Neospora...
Authors
Todd Atwood, Colleen Duncan, Kelly Patyk, Pauline Nol, Jack Rhyan, Matthew McCollum, Melissa McKinney, Andrew Ramey, Camila Cerqueira-Cezar, Oliver C H Kwok, Jitender Dubey, S.G. Hennager
Polar bears and sea ice habitat change Polar bears and sea ice habitat change
The polar bear (Ursus maritimus) is an obligate apex predator of Arctic sea ice and as such can be affected by climate warming-induced changes in the extent and composition of pack ice and its impacts on their seal prey. Sea ice declines have negatively impacted some polar bear subpopulations through reduced energy input because of loss of hunting habitats, higher energy costs due to...
Authors
George Durner, Todd Atwood
Uncertainties in forecasting the response of polar bears to global climate change Uncertainties in forecasting the response of polar bears to global climate change
Several sources of uncertainty affect how precisely the future status of polar bears (Ursus maritimus) can be forecasted. Foremost are unknowns about the future levels of global greenhouse gas emissions, which could range from an unabated increase to an aggressively mitigated reduction. Uncertainties also arise because different climate models project different amounts and rates of...
Authors
David Douglas, Todd Atwood
Ecological change drives a decline in mercury concentrations in southern Beaufort Sea polar bears Ecological change drives a decline in mercury concentrations in southern Beaufort Sea polar bears
We evaluated total mercury (THg) concentrations and trends in polar bears from the southern Beaufort Sea subpopulation from 2004 to 2011. Hair THg concentrations ranged widely among individuals from 0.6 to 13.3 μg g–1 dry weight (mean: 3.5 ± 0.2 μg g–1). Concentrations differed among sex and age classes: solitary adult females ≈ adult females with cubs ≈ subadults > adult males ≈...
Authors
Melissa McKinney, Todd Atwood, Sara Pedro, Elizabeth Peacock
Using tri-axial accelerometers to identify wild polar bear behaviors Using tri-axial accelerometers to identify wild polar bear behaviors
Tri-axial accelerometers have been used to remotely identify the behaviors of a wide range of taxa. Assigning behaviors to accelerometer data often involves the use of captive animals or surrogate species, as their accelerometer signatures are generally assumed to be similar to those of their wild counterparts. However, this has rarely been tested. Validated accelerometer data are needed...
Authors
Anthony Pagano, Karyn Rode, A. Cutting, M.A. Owen, S. Jensen, J.V. Ware, C.T. Robbins, George Durner, Todd Atwood, M.E. Obbard, K.R. Middel, G.W. Thiemann, T.M. Williams
Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land Collar temperature sensor data reveal long-term patterns in southern Beaufort Sea polar bear den distribution on pack ice and land
In response to a changing climate, many species alter habitat use. Polar bears Ursus maritimus in the southern Beaufort Sea have increasingly used land for maternal denning. To aid in detecting denning behavior, we developed an objective method to identify polar bear denning events using temperature sensor data collected by satellite-linked transmitters deployed on adult females between...
Authors
Jay Olson, Karyn Rode, Dennis Eggett, T. Smith, R. Wilson, George Durner, Anthony S. Fischbach, Todd Atwood, David C. Douglas
Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines Conservation status of polar bears (Ursus maritimus) in relation to projected sea-ice declines
Loss of Arctic sea ice owing to climate change is the primary threat to polar bears throughout their range. We evaluated the potential response of polar bears to sea-ice declines by (i) calculating generation length (GL) for the species, which determines the timeframe for conservation assessments; (ii) developing a standardized sea-ice metric representing important habitat; and (iii)...
Authors
Eric Regehr, Kristin Laidre, H. Akcakaya, Steven Amstrup, Todd Atwood, Nicholas J. Lunn, Martyn Obbard, Harry Stern, Gregory Thiemann, Øystein Wiig
Forecasting the relative influence of environmental and anthropogenic stressors on polar bears Forecasting the relative influence of environmental and anthropogenic stressors on polar bears
Effective conservation planning requires understanding and ranking threats to wildlife populations. We developed a Bayesian network model to evaluate the relative influence of environmental and anthropogenic stressors, and their mitigation, on the persistence of polar bears (Ursus maritimus). Overall sea ice conditions, affected by rising global temperatures, were the most influential...
Authors
Todd Atwood, Bruce Marcot, David Douglas, Steven Amstrup, Karyn Rode, George Durner, Jeffrey Bromaghin
Non-USGS Publications**
Algeo, T. P., D. Slate, R. M. Caron, T. C. Atwood, S. Recuenco, M. Ducey, R. B. Chipman, and M. Palace. 2017. Modeling raccoon (Procyon lotor) habitat connectivity to identify potential corridors for rabies spread. Tropical Medicine and Infectious Diseases 44. doi:10.3390/tropicalmed203044.
Atwood, T. C., E. Peacock, K. M. Lillie, R. R. Wilson, and S. Miller. 2015. Demographic composition and behavior of polar bears summering on shore in Alaska. USGS Administrative Report, 26 p.
Beasley, J. C., T. C. Atwood, M. E. Byrne, K. C. VerCauteren, S. R. Johnson, and O. E. Rhodes, Jr. 2015. A behaviorally-explicit approach for evaluating vaccine baits to mesopredators to control epizootics in fragmented landscapes. PLoS One 10:e0113206. doi:10.1371/journal.pone.0113206.
Anderson, A., S. A. Shwiff, R. B. Chipman, T. C. Atwood, T. Cozzens, F. Fillo, R. Hale, B. Hatch, J. Maki, O. E. Rhodes, Jr, E. E. Rees, C. E. Rupprecht, R. Tinline, K. C. VerCauteren, and D. Slate. 2014. Forecasting the spread of raccoon rabies using a purpose-specific group decision-making process. Human-Wildlife Interactions 8(1):130-138.
Slate, D., R. B. Chipman, T. P. Algeo, S. A. Mills, K. M. Nelson, C. K. Croson, E. J. Dubovi, R. W. Renshaw, K. C. VerCauteren, T. C. Atwood, S. Johnson, and C. E. Rupprecht. 2014. Safety and immunogenicity in the first field trial with ONRAB in raccoons in the United States. Journal of Wildlife Diseases 50(3):582-595. doi:10.7589/2013-08-207.
Kunkel, K. E., T. K. Ruth, T. C. Atwood, D. H. Pletscher, and M. G. Hornocker. 2013. Assessing the value of wolves and cougars as conservation surrogates by linking carnivore hunting success with landscape characteristics. Animal Conservation 16:32-40. doi:10.1111/j.1469-1795.2012.00568.x.
Beasley, J. D., W. S. Beatty, T. C. Atwood, S. Johnson, and O. E. Rhodes, Jr. 2012. A comparison of methods for estimating raccoon abundance: Implications for disease vaccination programs. Journal of Wildlife Management 76(6):1290-1297. doi:10.1002/jwmg.379.
Atwood, T. C. and S. W. Breck. 2012. Carnivores, Conflict, and Conservation: Defining the Landscape of Conflict. Pages 99-118 in F. I. Álvares and G. E. Mata, (eds.). Carnivores: Species, Conservation, and Management. Nova Publishers.
Atwood, T. C., T. L. Fry, and B. R. Leland. 2011. Partitioning of a limited resource by sympatric carnivores in the Chihuahuan Desert and the implications for disease transmission. Journal of Wildlife Management 75:1609-1615.
Atwood, T. C., J. K. Young, J. P. Beckmann, S. W. Breck, O. E. Rhodes, Jr, J. A. Fike, and K. D. Bristow. 2011. Modeling connectivity of black bears in a desert sky island archipelago. Biological Conservation 144(12):2851-2862. doi:10.1016/j.biocon.2011.08.002.
Fry, T. L., T. C. Atwood, and M. R. Dunbar. 2010. Utility of rhodamine B as a biomarker in raccoons. Human-Wildlife Interactions 4:275-282.
Atwood, T. C. and E. M. Gese. 2010. Importance of resource selection and social behaviour to partitioning of hostile space by sympatric canids. Journal of Mammalogy 91:490-499.
Atwood, T. C., T. J. DeLiberto, H. J. Smith, J. Stevenson, and K. C. VerCauteren. 2009. Raccoon spatial ecology related to cattle and bovine tuberculosis. Journal of Wildlife Management 73:647-654.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2009. Spatial decomposition of predation risk in a multiple-predator multiple-prey system. Journal of Wildlife Management 73:876-884.
Atwood, T. C. and E. M. Gese. 2008. Coyotes (Canis latrans) and recolonizing wolves (Canis lupus): Social rank mediates risk-conditional behaviour at ungulate carcasses. Animal Behaviour 75:753-762.
VerCauteren, K. C., T. C. Atwood, T. J. DeLiberto, H. J. Smith, J. Stevenson, T. Gidlewski, and B. V. Thomsen. 2008. Sentinel-based surveillance of coyotes to detect bovine tuberculosis in Michigan. Emerging Infectious Diseases 14:1862-1869.
Atwood, T. C., K. C. VerCauteren, T. J. DeLiberto, H. J. Smith, and J. Stevenson. 2007. Coyotes as a potential sentinel species to detect bovine tuberculosis (Mycobacterium bovis) infection in white-tailed deer in Michigan. Journal of Wildlife Management 71:1545-1554.
Atwood, T. C., E. M. Gese, and K. E. Kunkel. 2007. Comparative patterns of predation by cougars and recolonizing wolves. Journal of Wildlife Management 71:1098-1106.
Atwood, T. C. 2006. The influence of habitat patch attributes on coyote group size and interaction in a fragmented landscape. Canadian Journal of Zoology 84:80-87.
Atwood, T. C. 2006. Behavioral interactions between wolves, Canis lupus, and coyotes, Canis latrans, at ungulate carcasses in southwest Montana. Western North American Naturalist 66:390-394.
Atwood, T. C., H. P. Weeks, Jr., and T. M. Gehring. 2004. Spatial ecology of coyotes along a suburban-to-rural gradient. Journal of Wildlife Management 85:1000-1009.
Swihart, R. K., T. C. Atwood, J. R. Goheen, D. A. Scheiman, K. E. Munroe, and T. M. Gehring. 2003. Patch occupancy in North American mammals: Is patchiness in the eye of the beholder? . Journal of Biogeography 30:1259-1279.
Atwood, T. C. and H. P. Weeks, Jr.. 2003. Sex-specific patterns of mineral lick preference in white-tailed deer. Northeastern Naturalist 10:409-414.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Sex- and age-specific patterns of mineral lick use by white-tailed deer. American Midland Naturalist 148:289-296.
Atwood, T. C. and H. P. Weeks, Jr.. 2002. Facultative dyad formation in adult male coyotes. Northeastern Naturalist 9:353-358.
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
*Disclaimer: Listing outside positions with professional scientific organizations on this Staff Profile are for informational purposes only and do not constitute an endorsement of those professional scientific organizations or their activities by the USGS, Department of the Interior, or U.S. Government