Changes in proximate composition and somatic energy content for Pacific sand lance (Ammodytes hexapterus) from Kachemak Bay, Alaska relative to maturity and season

Mean dry-weight energy values of adult Pacific sand lance (Ammodytes hexapterus) peaked in spring and early summer (20.91 kJg⁻¹ for males, 21.08 kJg⁻¹ for females), then declined by about 25% during late summer and fall (15.91 kJg⁻¹ for males, 15.74 kJg⁻¹ for females). Late summer declines in energy density paralleled gonadal development. Gender differences in energy density (males