Distribution, abundance, and breeding activities of Southwestern Willow Flycatchers (Empidonax traillii extimus) on the San Dieguito River and upper San Luis Rey River, San Diego County, California—2025 data summary

Executive Summary 

We surveyed for Southwestern Willow Flycatchers (Empidonax traillii extimus; flycatcher) at the San Dieguito River and the upper San Luis Rey River in 2025. Surveys were completed at five locations: one along the San Dieguito River (San Dieguito [SD]), which was last surveyed in 2016, and four along the upper San Luis Rey River, including three downstream from Lake Henshaw that have been surveyed annually since 2015 (Rey River Ranch [RRR], Cleveland National Forest [CNF], Vista Irrigation District [VID]), and one upstream at VID Lake Henshaw (VLH) that has been surveyed annually since 2018. There was a minimum of 57 territorial flycatchers (22 male, 35 female) and 3 transient flycatchers of unknown subspecies detected at 1 location (VLH). In total, 37 territories were established, containing 35 pairs (20 males and 35 females) and 2 male flycatchers of undetermined breeding status. Of the 35 pairs, 12 were monogamous pairings, and 23 were polygynous pairings consisting of 3 males each pairing with 2 different females [6 pairs], 3 males each pairing with 3 different females [9 pairs], and 2 males each pairing with 4 different females [8 pairs]).

No territorial flycatchers were detected downstream from Lake Henshaw or along the San Dieguito River. Brown-headed Cowbirds (Molothrus ater; cowbird) were detected at all five survey locations. No banded flycatchers were detected during surveys.

Flycatchers used only one habitat type at VLH, mixed willow riparian. All flycatcher locations were in habitat characterized as mixed willow riparian dominated by Goodding’s black willow (Salix gooddingii), and 93 percent were in habitat with greater than 95-percent native plant cover.

We monitored flycatcher nests at VLH to collect baseline data on nest success, productivity, and cowbird parasitism rate. There were 33 completed nests monitored in 26 territories; 10 were successful (30 percent). Of the 23 failed nests, 14 were depredated, 5 failed for unknown reasons, and 4 failed because of cowbird parasitism. There were 33 fledglings confirmed in monitored territories, yielding a seasonal productivity of 1.3 young/pair (33 young/26 monitored pairs). One additional fledgling was confirmed in an unmonitored territory during surveys at VLH. Based on 31 nests in which the contents were observed during the egg stage, 23 percent of nests in 2025 were parasitized. In two additional territories where nests were not located, adult flycatchers were observed feeding a cowbird fledgling.