Mark Haroldson
Mark Haroldson is an emeritus USGS Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 90
Evidence for density-dependent effects on body composition of a large omnivore in a changing Greater Yellowstone Ecosystem Evidence for density-dependent effects on body composition of a large omnivore in a changing Greater Yellowstone Ecosystem
Understanding the density-dependent processes that drive population demography in a changing world is critical in ecology, yet measuring performance–density relationships in long-lived mammalian species demands long-term data, limiting scientists' ability to observe such mechanisms. We tested performance–density relationships for an opportunistic omnivore, grizzly bears (Ursus arctos...
Authors
Andrea Corradini, Mark A. Haroldson, Francesca Cagnacci, Cecily M. Costello, Daniel D. Bjornlie, Daniel Thompson, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Frank T. van Manen
Grizzly bear movement models predict habitat use for nearby populations Grizzly bear movement models predict habitat use for nearby populations
Conservation planning and decision-making can be enhanced by ecological models that reliably transfer to times and places beyond those where models were developed. Transferrable models can be especially helpful for species of conservation concern, such as grizzly bears (Ursus arctos). Currently, only four grizzly bear populations remain in the contiguous United States. We evaluated...
Authors
Sarah Nelson Sells, Cecily M. Costello, Paul Lukacs, Frank T. van Manen, Mark A. Haroldson, Wayne Kasworm, Justin Tesiberg, Milan Vinks, Daniel D. Bjornlie
Enhancements to population monitoring of Yellowstone grizzly bears Enhancements to population monitoring of Yellowstone grizzly bears
In the Greater Yellowstone Ecosystem, counts of female grizzly bears (Ursus arctos) with cubs-of-the-year (females with cubs) from systematic aerial surveys and opportunistic ground sightings are combined with demographic data to derive annual population estimates. We addressed 2 limitations to the monitoring approach. As part of a rule set, a conservative distance of >30 km currently is...
Authors
Frank T. van Manen, Michael Ebinger, Cecily M. Costello, Daniel D. Bjornlie, Justin Clapp, Daniel Thompson, Mark A. Haroldson, Kevin L. Frey, Curtis Hendricks, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Hilary Cooley, Jennifer Fortin-Noreus, Pat Hnilicka, Daniel B. Tyers
Genetic architecture and evolution of color variation in American black bears Genetic architecture and evolution of color variation in American black bears
Color variation is a frequent evolutionary substrate for camouflage in small mammals, but the underlying genetics and evolutionary forces that drive color variation in natural populations of large mammals are mostly unexplained. The American black bear, Ursus americanus (U. americanus), exhibits a range of colors including the cinnamon morph, which has a similar color to the brown bear...
Authors
E. Puckett, I. S. Davis, D. C. Harper, K. Wakamatsu, G. Battu, J. L. Belant, D. E. Beyer, C. Carpenter, A. P. Crupi, M. Davidson, C. S. DePerno, N. Forman, N. L. Fowler, D. L. Garshelis, N. Gould, K. Gunther, Mark A. Haroldson, S. Ito, David. M Kocka, C. Lackey, R. Leahy, C. Lee-Roney, T. Lewis, A. Lutto, K. McGowan, C. Olfenbuttel, M. Orlando, A. Platt, M. D. Pollard, M. Ramaker, Heather Reich, Jaime L. Sajecki, S. K. Sell, J. Strules, S. Thompson, Frank T. van Manen, Craig Whitman, R. Williamson, F. Winslow, C. B. Kaelin, M. S. Marks, G. S. Barsh
Agent-based models for collective animal movement: Proximity-induced state switching Agent-based models for collective animal movement: Proximity-induced state switching
Animal movement is a complex phenomenon where individual movement patterns can be influenced by a variety of factors including the animal’s current activity, available terrain and habitat, and locations of other animals. Motivated by modeling grizzly bear movement in the Greater Yellowstone Ecosystem, this article presents an agent-based model represented in a state-space framework for...
Authors
Andrew B. Hoegh, Frank T. van Manen, Mark A. Haroldson
Physiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears. Physiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears.
Meat, fruit, seeds and other high-energy bear foods are often highly localized and briefly available and understanding which factors influence bear consumption of these foods is a common focus of bear conservation and ecology. However, the most common bear foods, graminoids and forbs, are more widespread but of lower quality. We poorly understand how herbage consumption impacts bear...
Authors
David A Christianson, Tyler H Coleman, Quint Doan, Mark A. Haroldson
A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem
The Yellowstone Ecosystem Subcommittee (YES) asked the Interagency Grizzly Bear Study Team (IGBST) to re-assess a technique used in annual population estimation and trend monitoring of grizzly bears in the Greater Yellowstone Ecosystem (GYE). This technique is referred to as the Chao2 approach and estimates the number of females with cubs-of-the-year (hereafter, females with cubs) and...
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Daniel D. Bjornlie, Justin Clapp, Daniel J. Thompson, Kevin L. Frey, Cecily M. Costello, Curtis Hendricks, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Hilary Cooley, Jennifer Fortin-Noreus, Pat Hnilicka, Daniel B. Tyers
Quantifying energetic costs and defining energy landscapes experienced by grizzly bears Quantifying energetic costs and defining energy landscapes experienced by grizzly bears
Animal movements are major determinants of energy expenditure and ultimately the cost–benefit of landscape use. Thus, we sought to understand those costs and how grizzly bears (Ursus arctos) move in mountainous landscapes. We trained captive grizzly bears to walk on a horizontal treadmill and up and down 10% and 20% slopes. The cost of moving upslope increased linearly with speed and...
Authors
Anthony M. Carnahan, Frank T. van Manen, Mark A. Haroldson, Gordon Stenhouse, Charles T. Robbins
Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears
1. Heat dissipation limit theory posits that energy available for growth and reproduction in endotherms is limited by their ability to dissipate heat. In mammals, endogenous heat production increases markedly during gestation and lactation, and thus female mammals may be subject to greater thermal constraints on energy expenditure than males. Such constraints likely have important...
Authors
Savannah A. Rogers, Charlie T. Robbins, Paul D. Mathewson, Anthony M. Carnahan, Frank T. van Manen, Mark A. Haroldson, Warren P. Porter, Taylor R. Rogers, Terrence Soule, Ryan A. Long
Yellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team Yellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team
This Annual Report summarizes results of grizzly bear (Ursus arctos) research and monitoring conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2020. The research and monitoring program is focused on population estimation and demographics, food monitoring, and habitat monitoring. This report also presents a summary of grizzly...
Science and Products
Filter Total Items: 90
Evidence for density-dependent effects on body composition of a large omnivore in a changing Greater Yellowstone Ecosystem Evidence for density-dependent effects on body composition of a large omnivore in a changing Greater Yellowstone Ecosystem
Understanding the density-dependent processes that drive population demography in a changing world is critical in ecology, yet measuring performance–density relationships in long-lived mammalian species demands long-term data, limiting scientists' ability to observe such mechanisms. We tested performance–density relationships for an opportunistic omnivore, grizzly bears (Ursus arctos...
Authors
Andrea Corradini, Mark A. Haroldson, Francesca Cagnacci, Cecily M. Costello, Daniel D. Bjornlie, Daniel Thompson, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Frank T. van Manen
Grizzly bear movement models predict habitat use for nearby populations Grizzly bear movement models predict habitat use for nearby populations
Conservation planning and decision-making can be enhanced by ecological models that reliably transfer to times and places beyond those where models were developed. Transferrable models can be especially helpful for species of conservation concern, such as grizzly bears (Ursus arctos). Currently, only four grizzly bear populations remain in the contiguous United States. We evaluated...
Authors
Sarah Nelson Sells, Cecily M. Costello, Paul Lukacs, Frank T. van Manen, Mark A. Haroldson, Wayne Kasworm, Justin Tesiberg, Milan Vinks, Daniel D. Bjornlie
Enhancements to population monitoring of Yellowstone grizzly bears Enhancements to population monitoring of Yellowstone grizzly bears
In the Greater Yellowstone Ecosystem, counts of female grizzly bears (Ursus arctos) with cubs-of-the-year (females with cubs) from systematic aerial surveys and opportunistic ground sightings are combined with demographic data to derive annual population estimates. We addressed 2 limitations to the monitoring approach. As part of a rule set, a conservative distance of >30 km currently is...
Authors
Frank T. van Manen, Michael Ebinger, Cecily M. Costello, Daniel D. Bjornlie, Justin Clapp, Daniel Thompson, Mark A. Haroldson, Kevin L. Frey, Curtis Hendricks, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Hilary Cooley, Jennifer Fortin-Noreus, Pat Hnilicka, Daniel B. Tyers
Genetic architecture and evolution of color variation in American black bears Genetic architecture and evolution of color variation in American black bears
Color variation is a frequent evolutionary substrate for camouflage in small mammals, but the underlying genetics and evolutionary forces that drive color variation in natural populations of large mammals are mostly unexplained. The American black bear, Ursus americanus (U. americanus), exhibits a range of colors including the cinnamon morph, which has a similar color to the brown bear...
Authors
E. Puckett, I. S. Davis, D. C. Harper, K. Wakamatsu, G. Battu, J. L. Belant, D. E. Beyer, C. Carpenter, A. P. Crupi, M. Davidson, C. S. DePerno, N. Forman, N. L. Fowler, D. L. Garshelis, N. Gould, K. Gunther, Mark A. Haroldson, S. Ito, David. M Kocka, C. Lackey, R. Leahy, C. Lee-Roney, T. Lewis, A. Lutto, K. McGowan, C. Olfenbuttel, M. Orlando, A. Platt, M. D. Pollard, M. Ramaker, Heather Reich, Jaime L. Sajecki, S. K. Sell, J. Strules, S. Thompson, Frank T. van Manen, Craig Whitman, R. Williamson, F. Winslow, C. B. Kaelin, M. S. Marks, G. S. Barsh
Agent-based models for collective animal movement: Proximity-induced state switching Agent-based models for collective animal movement: Proximity-induced state switching
Animal movement is a complex phenomenon where individual movement patterns can be influenced by a variety of factors including the animal’s current activity, available terrain and habitat, and locations of other animals. Motivated by modeling grizzly bear movement in the Greater Yellowstone Ecosystem, this article presents an agent-based model represented in a state-space framework for...
Authors
Andrew B. Hoegh, Frank T. van Manen, Mark A. Haroldson
Physiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears. Physiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears.
Meat, fruit, seeds and other high-energy bear foods are often highly localized and briefly available and understanding which factors influence bear consumption of these foods is a common focus of bear conservation and ecology. However, the most common bear foods, graminoids and forbs, are more widespread but of lower quality. We poorly understand how herbage consumption impacts bear...
Authors
David A Christianson, Tyler H Coleman, Quint Doan, Mark A. Haroldson
A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem
The Yellowstone Ecosystem Subcommittee (YES) asked the Interagency Grizzly Bear Study Team (IGBST) to re-assess a technique used in annual population estimation and trend monitoring of grizzly bears in the Greater Yellowstone Ecosystem (GYE). This technique is referred to as the Chao2 approach and estimates the number of females with cubs-of-the-year (hereafter, females with cubs) and...
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Daniel D. Bjornlie, Justin Clapp, Daniel J. Thompson, Kevin L. Frey, Cecily M. Costello, Curtis Hendricks, Jeremy M. Nicholson, Kerry A. Gunther, Katharine R. Wilmot, Hilary Cooley, Jennifer Fortin-Noreus, Pat Hnilicka, Daniel B. Tyers
Quantifying energetic costs and defining energy landscapes experienced by grizzly bears Quantifying energetic costs and defining energy landscapes experienced by grizzly bears
Animal movements are major determinants of energy expenditure and ultimately the cost–benefit of landscape use. Thus, we sought to understand those costs and how grizzly bears (Ursus arctos) move in mountainous landscapes. We trained captive grizzly bears to walk on a horizontal treadmill and up and down 10% and 20% slopes. The cost of moving upslope increased linearly with speed and...
Authors
Anthony M. Carnahan, Frank T. van Manen, Mark A. Haroldson, Gordon Stenhouse, Charles T. Robbins
Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears
1. Heat dissipation limit theory posits that energy available for growth and reproduction in endotherms is limited by their ability to dissipate heat. In mammals, endogenous heat production increases markedly during gestation and lactation, and thus female mammals may be subject to greater thermal constraints on energy expenditure than males. Such constraints likely have important...
Authors
Savannah A. Rogers, Charlie T. Robbins, Paul D. Mathewson, Anthony M. Carnahan, Frank T. van Manen, Mark A. Haroldson, Warren P. Porter, Taylor R. Rogers, Terrence Soule, Ryan A. Long
Yellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team Yellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team
This Annual Report summarizes results of grizzly bear (Ursus arctos) research and monitoring conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2020. The research and monitoring program is focused on population estimation and demographics, food monitoring, and habitat monitoring. This report also presents a summary of grizzly...