Mark Haroldson
Mark Haroldson is an emeritus USGS Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 89
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013 Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the...
Authors
Kerry Gunther, Rebecca Shoemaker, Kevin Frey, Mark Haroldson, Steven Cain, Frank van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible...
Authors
Frank van Manen, Cecily M. Costello, Mark Haroldson, Daniel D. Bjornlie, Michael Ebinger, Kerry Gunther, Mary Mahalovich, Daniel Thompson, Megan Higgs, Kathryn Irvine, Kristin Legg, Daniel Tyers, Lisa Landenburger, Steven Cain, Kevin Frey, Bryan Aber, Charles Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the...
Authors
Megan D. Higgs, William Link, Gary White, Mark Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles Schwartz, Mark Haroldson, Kerry Gunther, Charles Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and...
Authors
Charles Schwartz, Jennifer K. Fortin, Justin Teisberg, Mark Haroldson, Christopher Servheen, Charles Robbins, Frank van Manen
Impacts of rural development on Yellowstone wildlife: linking grizzly bear Ursus arctos demographics with projected residential growth Impacts of rural development on Yellowstone wildlife: linking grizzly bear Ursus arctos demographics with projected residential growth
Exurban development is consuming wildlife habitat within the Greater Yellowstone Ecosystem with potential consequences to the long-term conservation of grizzly bears Ursus arctos. We assessed the impacts of alternative future land-use scenarios by linking an existing regression-based simulation model predicting rural development with a spatially explicit model that predicted bear...
Authors
Charles Schwartz, Patricia Gude, Lisa Landenburger, Mark Haroldson, Shannon Podruzny
Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States
Population fragmentation compromises population viability, reduces a species ability to respond to climate change, and ultimately may reduce biodiversity. We studied the current state and potential causes of fragmentation in grizzly bears over approximately 1,000,000 km 2 of western Canada, the northern United States (US), and southeast Alaska. We compiled much of our data from projects...
Authors
M.F. Proctor, David Paetkau, B. McLellan, G.B. Stenhouse, K.C. Kendall, R.D. Mace, W.F. Kasworm, C. Servheen, C.L. Lausen, M.L. Gibeau, W.L. Wakkinen, M.A. Haroldson, G. Mowat, C.D. Apps, L.M. Ciarniello, R.M.R. Barclay, M.S. Boyce, C.C. Schwartz, C. Strobeck
Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana
We estimated grizzly bear (Ursus arctos) population vital rates and trend for the Northern Continental Divide Ecosystem (NCDE), Montana, between 2004 and 2009 by following radio-collared females and observing their fate and reproductive performance. Our estimates of dependent cub and yearling survival were 0.612 (95% CI = 0.300–0.818) and 0.682 (95% CI = 0.258–0.898). Our estimates of...
Authors
R.D. Mace, D.W. Carney, T. Chilton-Radandt, S.A. Courville, M.A. Haroldson, R.B. Harris, J. Jonkel, B. McLellan, M. Madel, T.L. Manley, C.C. Schwartz, C. Servheen, G. Stenhouse, J.S. Waller, E. Wenum
Cougar survival and source-sink structure on Greater Yellowstone's Northern Range Cougar survival and source-sink structure on Greater Yellowstone's Northern Range
We studied survival and causes of mortality of radiocollared cougars (Puma concolor) on the Greater Yellowstone Northern Range (GYNR) prior to (1987–1994) and after wolf (Canis lupus) reintroduction (1998–2005) and evaluated temporal, spatial, and environmental factors that explain variation in adult, subadult, and kitten survival. Using Program MARK and multimodel inference, we modeled...
Authors
T.K. Ruth, M.A. Haroldson, K.M. Murphy, P.C. Buotte, M.G. Hornocker, H.B. Quigley
Study design and sampling intensity for demographic analyses of bear populations Study design and sampling intensity for demographic analyses of bear populations
The rate of population change through time (??) is a fundamental element of a wildlife population's conservation status, yet estimating it with acceptable precision for bears is difficult. For studies that follow known (usually marked) bears, ?? can be estimated during some defined time by applying either life-table or matrix projection methods to estimates of individual vital rates...
Authors
R.B. Harris, C.C. Schwartz, R.D. Mace, M.A. Haroldson
Hazards affecting grizzly bear survival in the Greater Yellowstone Ecosystem Hazards affecting grizzly bear survival in the Greater Yellowstone Ecosystem
During the past 2 decades, the grizzly bear (Ursus arctos) population in the Greater Yellowstone Ecosystem (GYE) has increased in numbers and expanded its range. Early efforts to model grizzly bear mortality were principally focused within the United States Fish and Wildlife Service Grizzly Bear Recovery Zone, which currently represents only about 61% of known bear distribution in the...
Authors
Charles Schwartz, Mark Haroldson, Gary C. White
Science and Products
Filter Total Items: 89
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013 Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the...
Authors
Kerry Gunther, Rebecca Shoemaker, Kevin Frey, Mark Haroldson, Steven Cain, Frank van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible...
Authors
Frank van Manen, Cecily M. Costello, Mark Haroldson, Daniel D. Bjornlie, Michael Ebinger, Kerry Gunther, Mary Mahalovich, Daniel Thompson, Megan Higgs, Kathryn Irvine, Kristin Legg, Daniel Tyers, Lisa Landenburger, Steven Cain, Kevin Frey, Bryan Aber, Charles Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the...
Authors
Megan D. Higgs, William Link, Gary White, Mark Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles Schwartz, Mark Haroldson, Kerry Gunther, Charles Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and...
Authors
Charles Schwartz, Jennifer K. Fortin, Justin Teisberg, Mark Haroldson, Christopher Servheen, Charles Robbins, Frank van Manen
Impacts of rural development on Yellowstone wildlife: linking grizzly bear Ursus arctos demographics with projected residential growth Impacts of rural development on Yellowstone wildlife: linking grizzly bear Ursus arctos demographics with projected residential growth
Exurban development is consuming wildlife habitat within the Greater Yellowstone Ecosystem with potential consequences to the long-term conservation of grizzly bears Ursus arctos. We assessed the impacts of alternative future land-use scenarios by linking an existing regression-based simulation model predicting rural development with a spatially explicit model that predicted bear...
Authors
Charles Schwartz, Patricia Gude, Lisa Landenburger, Mark Haroldson, Shannon Podruzny
Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States
Population fragmentation compromises population viability, reduces a species ability to respond to climate change, and ultimately may reduce biodiversity. We studied the current state and potential causes of fragmentation in grizzly bears over approximately 1,000,000 km 2 of western Canada, the northern United States (US), and southeast Alaska. We compiled much of our data from projects...
Authors
M.F. Proctor, David Paetkau, B. McLellan, G.B. Stenhouse, K.C. Kendall, R.D. Mace, W.F. Kasworm, C. Servheen, C.L. Lausen, M.L. Gibeau, W.L. Wakkinen, M.A. Haroldson, G. Mowat, C.D. Apps, L.M. Ciarniello, R.M.R. Barclay, M.S. Boyce, C.C. Schwartz, C. Strobeck
Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana
We estimated grizzly bear (Ursus arctos) population vital rates and trend for the Northern Continental Divide Ecosystem (NCDE), Montana, between 2004 and 2009 by following radio-collared females and observing their fate and reproductive performance. Our estimates of dependent cub and yearling survival were 0.612 (95% CI = 0.300–0.818) and 0.682 (95% CI = 0.258–0.898). Our estimates of...
Authors
R.D. Mace, D.W. Carney, T. Chilton-Radandt, S.A. Courville, M.A. Haroldson, R.B. Harris, J. Jonkel, B. McLellan, M. Madel, T.L. Manley, C.C. Schwartz, C. Servheen, G. Stenhouse, J.S. Waller, E. Wenum
Cougar survival and source-sink structure on Greater Yellowstone's Northern Range Cougar survival and source-sink structure on Greater Yellowstone's Northern Range
We studied survival and causes of mortality of radiocollared cougars (Puma concolor) on the Greater Yellowstone Northern Range (GYNR) prior to (1987–1994) and after wolf (Canis lupus) reintroduction (1998–2005) and evaluated temporal, spatial, and environmental factors that explain variation in adult, subadult, and kitten survival. Using Program MARK and multimodel inference, we modeled...
Authors
T.K. Ruth, M.A. Haroldson, K.M. Murphy, P.C. Buotte, M.G. Hornocker, H.B. Quigley
Study design and sampling intensity for demographic analyses of bear populations Study design and sampling intensity for demographic analyses of bear populations
The rate of population change through time (??) is a fundamental element of a wildlife population's conservation status, yet estimating it with acceptable precision for bears is difficult. For studies that follow known (usually marked) bears, ?? can be estimated during some defined time by applying either life-table or matrix projection methods to estimates of individual vital rates...
Authors
R.B. Harris, C.C. Schwartz, R.D. Mace, M.A. Haroldson
Hazards affecting grizzly bear survival in the Greater Yellowstone Ecosystem Hazards affecting grizzly bear survival in the Greater Yellowstone Ecosystem
During the past 2 decades, the grizzly bear (Ursus arctos) population in the Greater Yellowstone Ecosystem (GYE) has increased in numbers and expanded its range. Early efforts to model grizzly bear mortality were principally focused within the United States Fish and Wildlife Service Grizzly Bear Recovery Zone, which currently represents only about 61% of known bear distribution in the...
Authors
Charles Schwartz, Mark Haroldson, Gary C. White