Closure of the Isthmus of Panama: The near-shore marine record of Costa Rica and western Panama

The final closure of the Isthmus of Panama at ∼3.5 Ma divided the American tropical ocean into two separate and different oceanographic regions. Consequences for the marine biota were profound, but, hitherto, correlation of the Pacific and Caribbean coastal sections has not been precise enough to track biologic patterns. We present here a correlation of 31 sections from the Pacific and Caribbean coasts of Costa Rica and western Panama. Using calcareous nannofossils and planktonic foraminifera at both the tops and bottoms of each formation, we estimate that the Caribbean section ranges from 8.2 Ma to 1.7 Ma; and the Pacific sequence, from 3.6 Ma to