Neal Woodman, Ph.D. (Former Employee)
Science and Products
Rafinesque’s Mammals
The Challenge: Taxonomic nomenclature relies, in part, upon an accurate taxonomic history in order to establish the correct name for a taxon. Constantine S. Rafinesque (1783–1840), was a knowledgeable North American natural historian who was is responsible for describing and naming such iconic American mammals as the mule deer [Odocoileus hemionus (Rafinesque, 1817)] and the white-footed mouse...
Survival of the Least Fit: Incidence of Physical Trauma in a Wild Mammal Community
The Challenge: It has been generally considered that a severe injury to a wild mammal that seemingly limits its ability to forage for food or escape predators will almost certainly lead to that individual’s demise. Inspection of skeletons of wild caught small mammals, however, has revealed a surprising number of individuals with healed fractures of the skeletal bones―including the primary...
How Mammals Move: Locomotory Function in the Soricidae
The Challenge: The postcranial skeletons of mammals exhibit tremendous variation in form that partly relates to phylogeny (who a particular species is related to) and partly to locomotory function (how that species moves through its environment). Understanding the contributions of these two factors is important because phylogenetic characters assist in working out evolutionary relationships...
Biodiversity of North American Mammals
The Challenge: Despite more than a century and a half of study, accurate understanding of the diversity North American mammalian species and the distribution of those species remains unrefined. Yet this understanding is essential for determining the conservation status of species, for mapping out potential disease reservoirs, and for understanding the response of species to habitat perturbation...
Diversity and Biogeography of Treeshrews
The Challenge: Treeshrews (order Scandentia) are small-bodied mammals endemic to South and Southeast Asia. Since it was first described in 1820, the Common Treeshrew (Tupaia glis) has had a complex taxonomic history that has led to widely variable estimates of diversity, misidentification of populations, and general confusion regarding it and closely related species. One result is that T. glis has...
What Ancient Egyptian Shrew Mummies Reveal About Small Mammal Responses to Climate Change
The Challenge: Ancient Egyptians mummified animals for a variety of reasons, not the least of which was as votive offerings to certain deities. Among the six species of shrews that have been identified as mummies, one is now extinct, one is no longer occurs in Egypt, and the remaining four have more restricted distributions in the country. One of the latter species also exhibits significantly...
Filter Total Items: 76
Records of fleas (Siphonaptera) from Delaware
We present an annotated checklist of fleas (Siphonaptera) known to occur in the state of Delaware based on an examination of Siphonaptera collections at the University of Delaware and the Carnegie Museum of Natural History, as well as new specimens of fleas we collected from wildlife, other hosts, and tick flags. We review published records and compile them herein with our new records...
Authors
A.C Kennedy, W.S. Winter, A.L. Gardner, Neal Woodman, S.A. Shifflett, S. Redus, J.R. Newcomer, R. P. Eckerlin
Two new species of small-eared shrews of the Genus Cryptotis Pomel, 1848, from the Colombian Andes (Mammalia: Eulipotyphla: Soricidae)
Shrews (Mammalia: Eulipotyphla: Soricidae) reach the southern limit of their New World distribution in the Andes and eastern coastal highlands of northern South America. South of Honduras, the family is represented only by species of the genus Cryptotis Pomel, 1848. In South America, soricids are restricted to moist, high-elevation environments above 1000 m, and their distribution...
Authors
Neal Woodman
Skeletal indicators of locomotor adaptations in shrews
The Soricidae (Mammalia: Eulypotyphla) comprises more than 450 species inhabiting a variety of habitats on five continents. As a family, shrews employ a variety of locomotor modes that incorporate ambulatory, fossorial, aquatic, and scansorial behaviors, illustrating an ability to exploit a variety of natural substrates and their associated resources. In this study, the association of...
Authors
Neal Woodman
Environmental implications of Ptolemaic Period rodents and shrews from the Sacred Falcon Necropolis at Quesna, Egypt (Mammalia: Muridae and Soricidae)
BackgroundAssemblages of mummified and preserved animals in necropoleis of Ptolemaic Period Egypt (ca. 332–30 BC) document some aspects of the ceremonial and religious practices of the ancient Egyptians, but study of these animal remains can also provide insight into the local environments in which the animals and humans lived.ResultsExcavations of the Sacred Falcon Necropolis at Quesna...
Authors
Neal Woodman, Salima Ikram, Joanne Rowland
Taxonomic boundaries in Lesser Treeshrews (Scandentia, Tupaiidae: Tupaia minor)
The Lesser Treeshrew, Tupaia minor Günther, 1876, is a small mammal from Southeast Asia with four currently recognized subspecies: T. m. minor from Borneo; T. m. malaccana from the Malay Peninsula; T. m. humeralis from Sumatra; and T. m. sincepis from Singkep Island and Lingga Island. A fifth subspecies, T. m. caedis, was previously synonymized with T. m. minor; it was thought to occur...
Authors
M. M. Juman, Neal Woodman, A. Miller-Murthy, Link E. Olson, E. J. Sargis
Whence Orycteropus? The correct authorship and date for the generic name of the aardvark (Mammalia, Tubulidentata, Orycteropodidae)
All else being equal, the principle of priority in zoological taxonomic nomenclature gives precedence to the earliest name for a particular taxon. Determining the origin of some late eighteenth- and early nineteenth-century taxonomic names, however, can be vexing, particularly when the history of a name was never completely documented in contemporary synonymies. The authorship and date...
Authors
Neal Woodman
Ecogeographic variation and taxonomic boundaries in Large Treeshrews (Scandentia, Tupaiidae: Tupaia tana Raffles, 1821) from Southeast Asia
The Large Treeshrew, Tupaia tana Raffles, 1821, is a small mammal (~205 g) from Southeast Asia with a complicated taxonomic history. Currently, 15 subspecies are recognized from Borneo, Sumatra, and smaller islands, and many were originally differentiated based on minor pelage differences and small sample sizes. We explored intraspecific variation in T. tana using quantitative...
Authors
Maya M. Juman, Neal Woodman, Link E. Olson, Eric J. Sargis
A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger–Volta interfluvium in mammalian biogeography
Tree hyraxes (Dendrohyrax) are one of only three genera currently recognized in Procaviidae, the only extant family in the mammalian order Hyracoidea. Their taxonomy and natural history have received little attention in recent decades. All tree hyrax populations of Guineo-Congolian forests of Africa are currently treated as a single species, Dendrohyrax dorsalis, the western tree hyrax...
Authors
John F. Oates, Neal Woodman, Philippe Gaubert, Eric J. Sargis, Edward D. Wiafe, Emilie Lecompte, Francoise Dowsett-Lemaire, Robert J. Dowsett, Sery Gonedele Bi, Rachel A. Ikemeh, Chabi A.M.S. Djagoun, Louise Tomsett, Simon K. Bearder
The limitations of external measurements for aging small mammals: The cautionary example of the Lesser Treeshrew (Scandentia: Tupaiidae: Tupaia minor Günther, 1876)
Age is a basic demographic characteristic vital to studies of mammalian social organization, population dynamics, and behavior. To eliminate potentially confounding ontogenetic variation, morphological comparisons among populations of mammals typically are limited to mature individuals (i.e., those assumed to have ceased most somatic growth). In our morphometric studies of treeshrews...
Authors
Neal Woodman, Ananth Miller-Murthy, Link E. Olson, Eric J. Sargis
A new addition to the embalmed fauna of ancient Egypt: Güldenstaedt’s White-toothed Shrew, Crocidura gueldenstaedtii (Pallas, 1811) (Mammalia: Eulipotyphla: Soricidae)
The Falcon Necropolis at Quesna in the Nile Delta of Egypt is considered to have been founded by the priest Djedhor, the Saviour, of Athribis (Tell Atrib in modern Benha) at the beginning of the Ptolemaic Period. Recent excavations here have revealed abundant avian remains from mummies dedicated to the ancient Egyptian god Horus Khenty-Khety. Among the few mammal remains from the site...
Authors
Neal Woodman, Salima Ikram, Joanne Rowland
The relevance of a type locality: The case of Mephitis interrupta Rafinesque, 1820 (Carnivora: Mephitidae)
For more than 130 years, the type locality of the Plains Spotted Skunk, Spilogale putorius interrupta (Rafinesque, 1820) has been accepted to be along the upper Missouri River. The species’ description was based on a specimen observed by Constantine S. Rafinesque during his 1818 exploration of the Ohio River Valley, but Rafinesque never ventured into the animal’s geographic range west of...
Authors
Neal Woodman, Adam W. Ferguson
The green mole, Astromycter prasinatus T. M. Harris, 1825 (Mammalia: Eulipotyphla: Talpidae): An origin story
Thaddeus William Harris described the green mole of Maine, Condylura prasinata (Mammalia: Eulipotyphla: Talpidae), in the July 1825 issue of the Boston Journal of Philosophy and the Arts, and this was considered the original description of the species throughout most of the nineteenth century. In the early twentieth century, taxonomists began instead to cite an earlier notice in the June...
Authors
Neal Woodman
Science and Products
Rafinesque’s Mammals
The Challenge: Taxonomic nomenclature relies, in part, upon an accurate taxonomic history in order to establish the correct name for a taxon. Constantine S. Rafinesque (1783–1840), was a knowledgeable North American natural historian who was is responsible for describing and naming such iconic American mammals as the mule deer [Odocoileus hemionus (Rafinesque, 1817)] and the white-footed mouse...
Survival of the Least Fit: Incidence of Physical Trauma in a Wild Mammal Community
The Challenge: It has been generally considered that a severe injury to a wild mammal that seemingly limits its ability to forage for food or escape predators will almost certainly lead to that individual’s demise. Inspection of skeletons of wild caught small mammals, however, has revealed a surprising number of individuals with healed fractures of the skeletal bones―including the primary...
How Mammals Move: Locomotory Function in the Soricidae
The Challenge: The postcranial skeletons of mammals exhibit tremendous variation in form that partly relates to phylogeny (who a particular species is related to) and partly to locomotory function (how that species moves through its environment). Understanding the contributions of these two factors is important because phylogenetic characters assist in working out evolutionary relationships...
Biodiversity of North American Mammals
The Challenge: Despite more than a century and a half of study, accurate understanding of the diversity North American mammalian species and the distribution of those species remains unrefined. Yet this understanding is essential for determining the conservation status of species, for mapping out potential disease reservoirs, and for understanding the response of species to habitat perturbation...
Diversity and Biogeography of Treeshrews
The Challenge: Treeshrews (order Scandentia) are small-bodied mammals endemic to South and Southeast Asia. Since it was first described in 1820, the Common Treeshrew (Tupaia glis) has had a complex taxonomic history that has led to widely variable estimates of diversity, misidentification of populations, and general confusion regarding it and closely related species. One result is that T. glis has...
What Ancient Egyptian Shrew Mummies Reveal About Small Mammal Responses to Climate Change
The Challenge: Ancient Egyptians mummified animals for a variety of reasons, not the least of which was as votive offerings to certain deities. Among the six species of shrews that have been identified as mummies, one is now extinct, one is no longer occurs in Egypt, and the remaining four have more restricted distributions in the country. One of the latter species also exhibits significantly...
Filter Total Items: 76
Records of fleas (Siphonaptera) from Delaware
We present an annotated checklist of fleas (Siphonaptera) known to occur in the state of Delaware based on an examination of Siphonaptera collections at the University of Delaware and the Carnegie Museum of Natural History, as well as new specimens of fleas we collected from wildlife, other hosts, and tick flags. We review published records and compile them herein with our new records...
Authors
A.C Kennedy, W.S. Winter, A.L. Gardner, Neal Woodman, S.A. Shifflett, S. Redus, J.R. Newcomer, R. P. Eckerlin
Two new species of small-eared shrews of the Genus Cryptotis Pomel, 1848, from the Colombian Andes (Mammalia: Eulipotyphla: Soricidae)
Shrews (Mammalia: Eulipotyphla: Soricidae) reach the southern limit of their New World distribution in the Andes and eastern coastal highlands of northern South America. South of Honduras, the family is represented only by species of the genus Cryptotis Pomel, 1848. In South America, soricids are restricted to moist, high-elevation environments above 1000 m, and their distribution...
Authors
Neal Woodman
Skeletal indicators of locomotor adaptations in shrews
The Soricidae (Mammalia: Eulypotyphla) comprises more than 450 species inhabiting a variety of habitats on five continents. As a family, shrews employ a variety of locomotor modes that incorporate ambulatory, fossorial, aquatic, and scansorial behaviors, illustrating an ability to exploit a variety of natural substrates and their associated resources. In this study, the association of...
Authors
Neal Woodman
Environmental implications of Ptolemaic Period rodents and shrews from the Sacred Falcon Necropolis at Quesna, Egypt (Mammalia: Muridae and Soricidae)
BackgroundAssemblages of mummified and preserved animals in necropoleis of Ptolemaic Period Egypt (ca. 332–30 BC) document some aspects of the ceremonial and religious practices of the ancient Egyptians, but study of these animal remains can also provide insight into the local environments in which the animals and humans lived.ResultsExcavations of the Sacred Falcon Necropolis at Quesna...
Authors
Neal Woodman, Salima Ikram, Joanne Rowland
Taxonomic boundaries in Lesser Treeshrews (Scandentia, Tupaiidae: Tupaia minor)
The Lesser Treeshrew, Tupaia minor Günther, 1876, is a small mammal from Southeast Asia with four currently recognized subspecies: T. m. minor from Borneo; T. m. malaccana from the Malay Peninsula; T. m. humeralis from Sumatra; and T. m. sincepis from Singkep Island and Lingga Island. A fifth subspecies, T. m. caedis, was previously synonymized with T. m. minor; it was thought to occur...
Authors
M. M. Juman, Neal Woodman, A. Miller-Murthy, Link E. Olson, E. J. Sargis
Whence Orycteropus? The correct authorship and date for the generic name of the aardvark (Mammalia, Tubulidentata, Orycteropodidae)
All else being equal, the principle of priority in zoological taxonomic nomenclature gives precedence to the earliest name for a particular taxon. Determining the origin of some late eighteenth- and early nineteenth-century taxonomic names, however, can be vexing, particularly when the history of a name was never completely documented in contemporary synonymies. The authorship and date...
Authors
Neal Woodman
Ecogeographic variation and taxonomic boundaries in Large Treeshrews (Scandentia, Tupaiidae: Tupaia tana Raffles, 1821) from Southeast Asia
The Large Treeshrew, Tupaia tana Raffles, 1821, is a small mammal (~205 g) from Southeast Asia with a complicated taxonomic history. Currently, 15 subspecies are recognized from Borneo, Sumatra, and smaller islands, and many were originally differentiated based on minor pelage differences and small sample sizes. We explored intraspecific variation in T. tana using quantitative...
Authors
Maya M. Juman, Neal Woodman, Link E. Olson, Eric J. Sargis
A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger–Volta interfluvium in mammalian biogeography
Tree hyraxes (Dendrohyrax) are one of only three genera currently recognized in Procaviidae, the only extant family in the mammalian order Hyracoidea. Their taxonomy and natural history have received little attention in recent decades. All tree hyrax populations of Guineo-Congolian forests of Africa are currently treated as a single species, Dendrohyrax dorsalis, the western tree hyrax...
Authors
John F. Oates, Neal Woodman, Philippe Gaubert, Eric J. Sargis, Edward D. Wiafe, Emilie Lecompte, Francoise Dowsett-Lemaire, Robert J. Dowsett, Sery Gonedele Bi, Rachel A. Ikemeh, Chabi A.M.S. Djagoun, Louise Tomsett, Simon K. Bearder
The limitations of external measurements for aging small mammals: The cautionary example of the Lesser Treeshrew (Scandentia: Tupaiidae: Tupaia minor Günther, 1876)
Age is a basic demographic characteristic vital to studies of mammalian social organization, population dynamics, and behavior. To eliminate potentially confounding ontogenetic variation, morphological comparisons among populations of mammals typically are limited to mature individuals (i.e., those assumed to have ceased most somatic growth). In our morphometric studies of treeshrews...
Authors
Neal Woodman, Ananth Miller-Murthy, Link E. Olson, Eric J. Sargis
A new addition to the embalmed fauna of ancient Egypt: Güldenstaedt’s White-toothed Shrew, Crocidura gueldenstaedtii (Pallas, 1811) (Mammalia: Eulipotyphla: Soricidae)
The Falcon Necropolis at Quesna in the Nile Delta of Egypt is considered to have been founded by the priest Djedhor, the Saviour, of Athribis (Tell Atrib in modern Benha) at the beginning of the Ptolemaic Period. Recent excavations here have revealed abundant avian remains from mummies dedicated to the ancient Egyptian god Horus Khenty-Khety. Among the few mammal remains from the site...
Authors
Neal Woodman, Salima Ikram, Joanne Rowland
The relevance of a type locality: The case of Mephitis interrupta Rafinesque, 1820 (Carnivora: Mephitidae)
For more than 130 years, the type locality of the Plains Spotted Skunk, Spilogale putorius interrupta (Rafinesque, 1820) has been accepted to be along the upper Missouri River. The species’ description was based on a specimen observed by Constantine S. Rafinesque during his 1818 exploration of the Ohio River Valley, but Rafinesque never ventured into the animal’s geographic range west of...
Authors
Neal Woodman, Adam W. Ferguson
The green mole, Astromycter prasinatus T. M. Harris, 1825 (Mammalia: Eulipotyphla: Talpidae): An origin story
Thaddeus William Harris described the green mole of Maine, Condylura prasinata (Mammalia: Eulipotyphla: Talpidae), in the July 1825 issue of the Boston Journal of Philosophy and the Arts, and this was considered the original description of the species throughout most of the nineteenth century. In the early twentieth century, taxonomists began instead to cite an earlier notice in the June...
Authors
Neal Woodman
*Disclaimer: Listing outside positions with professional scientific organizations on this Staff Profile are for informational purposes only and do not constitute an endorsement of those professional scientific organizations or their activities by the USGS, Department of the Interior, or U.S. Government