Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Education
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Science and Products
Interagency Grizzly Bear Study Team
The Interagency Grizzly Bear Study Team (IGBST) is an interdisciplinary group of scientists and biologists responsible for long-term monitoring and research efforts on grizzly bears in the Greater Yellowstone Ecosystem (GYE). The team was formed by the Department of the Interior (DOI) in 1973 as a direct result of controversy surrounding the closure of open pit garbage dumps within Yellowstone...
NOROCK Large Carnivore Research Program
NOROCK has substantial expertise in large carnivore research, primarily involving species listed as Threatened or Endangered. NOROCK’s Large Carnivore Research Program includes scientists from NOROCK’s Headquarters, West Glacier Field Station, and the Southern Appalachian Field Station. Studies are conducted in a wide variety of landscapes throughout the U.S., as well as international research...
Documented known and probable grizzly bear mortalities in the Greater Yellowstone Ecosystem, 2015-2021
We evaluate mortalities for population segments within the DMA by deriving estimates of total mortality for independent-age (equal to or less than 2 years old) females and independent-age males, including estimates of unknown/unreported mortalities based on Cherry et al. (2002). We then determine the total annual mortality rate for these segments as a percent of their respective population estimat
Detection histories of grizzly bears in Grand Teton National Park, 2014-2015
This dataset contains detection histories of grizzly bears in areas of Grand Teton National Park that are open for elk harvest as part of the Elk Reduction Program. Sampling followed a robust design analysis with 6 primary sampling periods of 4 weekly secondary sampling periods each during 2014-2015. In both years, primary sampling periods were in July-August (no elk hunt), September-October (no e
Canine distemper virus antibody titer results for grizzly bears and wolves in the Greater Yellowstone Ecosystem 1984-2014
We investigated the dynamics of canine distemper virus (CDV) in grizzly bears (Ursus arctos) and wolves (Canis lupus) of the Greater Yellowstone Ecosystem using serological data collected from 1984 to 2014. 565 sera samples were obtained from 425 unique grizzly bears (134 females and 291 males) from 1984 to 2014 and 319 sera samples were obtained from 285 unique wolves (130 females and 155 males)
Potential movement paths for male grizzly bear (Ursus arctos) dispersal between the Northern Continental Divide and Greater Yellowstone Ecosystems, 2000-2015
For several decades, grizzly bear (Ursus arctos) populations in the Greater Yellowstone Ecosystem (GYE) and the Northern Continental Divide Ecosystem (NCDE) have increased in numbers and range extent. Whereas the NCDE population is contiguous with grizzly bear populations in the Canadian Rocky Mountains, genetic evidence suggests the GYE population remains isolated. Recent analyses indicate the ef
Filter Total Items: 74
Enhancements to population monitoring of Yellowstone grizzly bears
In the Greater Yellowstone Ecosystem, counts of female grizzly bears (Ursus arctos) with cubs-of-the-year (females with cubs) from systematic aerial surveys and opportunistic ground sightings are combined with demographic data to derive annual population estimates. We addressed 2 limitations to the monitoring approach. As part of a rule set, a conservative distance of >30 km currently is used as a
Genetic architecture and evolution of color variation in American black bears
Color variation is a frequent evolutionary substrate for camouflage in small mammals, but the underlying genetics and evolutionary forces that drive color variation in natural populations of large mammals are mostly unexplained. The American black bear, Ursus americanus (U. americanus), exhibits a range of colors including the cinnamon morph, which has a similar color to the brown bear, U. arctos,
Agent-based models for collective animal movement: Proximity-induced state switching
Animal movement is a complex phenomenon where individual movement patterns can be influenced by a variety of factors including the animal’s current activity, available terrain and habitat, and locations of other animals. Motivated by modeling grizzly bear movement in the Greater Yellowstone Ecosystem, this article presents an agent-based model represented in a state-space framework for collective
Physiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears.
Meat, fruit, seeds and other high-energy bear foods are often highly localized and briefly available and understanding which factors influence bear consumption of these foods is a common focus of bear conservation and ecology. However, the most common bear foods, graminoids and forbs, are more widespread but of lower quality. We poorly understand how herbage consumption impacts bear physiology, su
A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem
The Yellowstone Ecosystem Subcommittee (YES) asked the Interagency Grizzly Bear Study Team (IGBST) to re-assess a technique used in annual population estimation and trend monitoring of grizzly bears in the Greater Yellowstone Ecosystem (GYE). This technique is referred to as the Chao2 approach and estimates the number of females with cubs-of-the-year (hereafter, females with cubs) and, in associat
Quantifying energetic costs and defining energy landscapes experienced by grizzly bears
Animal movements are major determinants of energy expenditure and ultimately the cost–benefit of landscape use. Thus, we sought to understand those costs and how grizzly bears (Ursus arctos) move in mountainous landscapes. We trained captive grizzly bears to walk on a horizontal treadmill and up and down 10% and 20% slopes. The cost of moving upslope increased linearly with speed and slope angle,
Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears
1. Heat dissipation limit theory posits that energy available for growth and reproduction in endotherms is limited by their ability to dissipate heat. In mammals, endogenous heat production increases markedly during gestation and lactation, and thus female mammals may be subject to greater thermal constraints on energy expenditure than males. Such constraints likely have important implications for
Yellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team
This Annual Report summarizes results of grizzly bear (Ursus arctos) research and monitoring conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2020. The research and monitoring program is focused on population estimation and demographics, food monitoring, and habitat monitoring. This report also presents a summary of grizzly bear managem
Brown Bear (Ursus arctos; North America)
This chapter comprises the following sections: names, taxonomy, subspecies and distribution, descriptive notes, habitat, movements and home range, activity patterns, feeding ecology, reproduction and growth, behavior, parasites and diseases, status in the wild, and status in captivity.
Whitebark pine cone production - 2020
No abstract available.
Potential for recreational restrictions to reduce grizzly bear–caused human injuries
In 2011, 2 hikers were killed by grizzly bears (Ursus arctos) in separate incidents on backcountry trails in Hayden Valley, Yellowstone National Park, USA (YNP). Hayden Valley provides prime habitat for grizzly bears and is known to have high densities of bears. During 1970–2017, 23% (10 of 44) of all backcountry grizzly bear–inflicted human injuries and fatalities in YNP occurred in the valley ev
Science and Products
- Science
Interagency Grizzly Bear Study Team
The Interagency Grizzly Bear Study Team (IGBST) is an interdisciplinary group of scientists and biologists responsible for long-term monitoring and research efforts on grizzly bears in the Greater Yellowstone Ecosystem (GYE). The team was formed by the Department of the Interior (DOI) in 1973 as a direct result of controversy surrounding the closure of open pit garbage dumps within Yellowstone...NOROCK Large Carnivore Research Program
NOROCK has substantial expertise in large carnivore research, primarily involving species listed as Threatened or Endangered. NOROCK’s Large Carnivore Research Program includes scientists from NOROCK’s Headquarters, West Glacier Field Station, and the Southern Appalachian Field Station. Studies are conducted in a wide variety of landscapes throughout the U.S., as well as international research... - Data
Documented known and probable grizzly bear mortalities in the Greater Yellowstone Ecosystem, 2015-2021
We evaluate mortalities for population segments within the DMA by deriving estimates of total mortality for independent-age (equal to or less than 2 years old) females and independent-age males, including estimates of unknown/unreported mortalities based on Cherry et al. (2002). We then determine the total annual mortality rate for these segments as a percent of their respective population estimatDetection histories of grizzly bears in Grand Teton National Park, 2014-2015
This dataset contains detection histories of grizzly bears in areas of Grand Teton National Park that are open for elk harvest as part of the Elk Reduction Program. Sampling followed a robust design analysis with 6 primary sampling periods of 4 weekly secondary sampling periods each during 2014-2015. In both years, primary sampling periods were in July-August (no elk hunt), September-October (no eCanine distemper virus antibody titer results for grizzly bears and wolves in the Greater Yellowstone Ecosystem 1984-2014
We investigated the dynamics of canine distemper virus (CDV) in grizzly bears (Ursus arctos) and wolves (Canis lupus) of the Greater Yellowstone Ecosystem using serological data collected from 1984 to 2014. 565 sera samples were obtained from 425 unique grizzly bears (134 females and 291 males) from 1984 to 2014 and 319 sera samples were obtained from 285 unique wolves (130 females and 155 males)Potential movement paths for male grizzly bear (Ursus arctos) dispersal between the Northern Continental Divide and Greater Yellowstone Ecosystems, 2000-2015
For several decades, grizzly bear (Ursus arctos) populations in the Greater Yellowstone Ecosystem (GYE) and the Northern Continental Divide Ecosystem (NCDE) have increased in numbers and range extent. Whereas the NCDE population is contiguous with grizzly bear populations in the Canadian Rocky Mountains, genetic evidence suggests the GYE population remains isolated. Recent analyses indicate the ef - Publications
Filter Total Items: 74
Enhancements to population monitoring of Yellowstone grizzly bears
In the Greater Yellowstone Ecosystem, counts of female grizzly bears (Ursus arctos) with cubs-of-the-year (females with cubs) from systematic aerial surveys and opportunistic ground sightings are combined with demographic data to derive annual population estimates. We addressed 2 limitations to the monitoring approach. As part of a rule set, a conservative distance of >30 km currently is used as aGenetic architecture and evolution of color variation in American black bears
Color variation is a frequent evolutionary substrate for camouflage in small mammals, but the underlying genetics and evolutionary forces that drive color variation in natural populations of large mammals are mostly unexplained. The American black bear, Ursus americanus (U. americanus), exhibits a range of colors including the cinnamon morph, which has a similar color to the brown bear, U. arctos,Agent-based models for collective animal movement: Proximity-induced state switching
Animal movement is a complex phenomenon where individual movement patterns can be influenced by a variety of factors including the animal’s current activity, available terrain and habitat, and locations of other animals. Motivated by modeling grizzly bear movement in the Greater Yellowstone Ecosystem, this article presents an agent-based model represented in a state-space framework for collectivePhysiological consequences of consuming low-energy foods: Herbivory coincides with a stress response in Yellowstone bears.
Meat, fruit, seeds and other high-energy bear foods are often highly localized and briefly available and understanding which factors influence bear consumption of these foods is a common focus of bear conservation and ecology. However, the most common bear foods, graminoids and forbs, are more widespread but of lower quality. We poorly understand how herbage consumption impacts bear physiology, suA reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem
The Yellowstone Ecosystem Subcommittee (YES) asked the Interagency Grizzly Bear Study Team (IGBST) to re-assess a technique used in annual population estimation and trend monitoring of grizzly bears in the Greater Yellowstone Ecosystem (GYE). This technique is referred to as the Chao2 approach and estimates the number of females with cubs-of-the-year (hereafter, females with cubs) and, in associatQuantifying energetic costs and defining energy landscapes experienced by grizzly bears
Animal movements are major determinants of energy expenditure and ultimately the cost–benefit of landscape use. Thus, we sought to understand those costs and how grizzly bears (Ursus arctos) move in mountainous landscapes. We trained captive grizzly bears to walk on a horizontal treadmill and up and down 10% and 20% slopes. The cost of moving upslope increased linearly with speed and slope angle,Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears
1. Heat dissipation limit theory posits that energy available for growth and reproduction in endotherms is limited by their ability to dissipate heat. In mammals, endogenous heat production increases markedly during gestation and lactation, and thus female mammals may be subject to greater thermal constraints on energy expenditure than males. Such constraints likely have important implications forYellowstone grizzly bear investigations 2020 - Annual report of the Interagency Grizzly Bear Study Team
This Annual Report summarizes results of grizzly bear (Ursus arctos) research and monitoring conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2020. The research and monitoring program is focused on population estimation and demographics, food monitoring, and habitat monitoring. This report also presents a summary of grizzly bear managemBrown Bear (Ursus arctos; North America)
This chapter comprises the following sections: names, taxonomy, subspecies and distribution, descriptive notes, habitat, movements and home range, activity patterns, feeding ecology, reproduction and growth, behavior, parasites and diseases, status in the wild, and status in captivity.Whitebark pine cone production - 2020
No abstract available.Potential for recreational restrictions to reduce grizzly bear–caused human injuries
In 2011, 2 hikers were killed by grizzly bears (Ursus arctos) in separate incidents on backcountry trails in Hayden Valley, Yellowstone National Park, USA (YNP). Hayden Valley provides prime habitat for grizzly bears and is known to have high densities of bears. During 1970–2017, 23% (10 of 44) of all backcountry grizzly bear–inflicted human injuries and fatalities in YNP occurred in the valley ev