Mark Haroldson
Biography
Education
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Research Interest
Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT. Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Science and Products
Interagency Grizzly Bear Study Team
The Interagency Grizzly Bear Study Team (IGBST) is an interdisciplinary group of scientists and biologists responsible for long-term monitoring and research efforts on grizzly bears in the Greater Yellowstone Ecosystem (GYE). The team was formed by the Department of the Interior (DOI) in 1973 as a direct result of controversy...
NOROCK Large Carnivore Research Program
NOROCK has substantial expertise in large carnivore research, primarily involving species listed as Threatened or Endangered. NOROCK’s Large Carnivore Research Program includes scientists from NOROCK’s Headquarters, West Glacier Field Station, and the Southern Appalachian Field Station. Studies are conducted in a wide variety of landscapes throughout the U.S., as well as international...
Agent-based models for collective animal movement: Proximity-induced state switching
Animal movement is a complex phenomenon where individual movement patterns can be influenced by a variety of factors including the animal’s current activity, available terrain and habitat, and locations of other animals. Motivated by modeling grizzly bear movement in the Greater Yellowstone Ecosystem, this article presents an agent-based model...
Hoegh, Andrew B.; van Manen, Frank T.; Haroldson, Mark A. A reassessment of Chao2 estimates for population monitoring of grizzly bears in the Greater Yellowstone Ecosystem
The Yellowstone Ecosystem Subcommittee (YES) asked the Interagency Grizzly Bear Study Team (IGBST) to re-assess a technique used in annual population estimation and trend monitoring of grizzly bears in the Greater Yellowstone Ecosystem (GYE). This technique is referred to as the Chao2 approach and estimates the number of females with cubs-of-the-...
van Manen, Frank T.; Ebinger, Michael R.; Haroldson, Mark A.; Bjornlie, Daniel D.; Clapp, Justin; Thompson, Daniel J.; Frey, Kevin L.; Costello, Cecily M.; Hendricks, Curtis; Nicholson, Jeremy M.; Gunther, Kerry A.; Wilmot, Katharine R.; Cooley, Hilary; Fortin-Noreus, Jennifer; Hnilicka, Pat; Tyers, Daniel B. Quantifying energetic costs and defining energy landscapes experienced by grizzly bears
Animal movements are major determinants of energy expenditure and ultimately the cost–benefit of landscape use. Thus, we sought to understand those costs and how grizzly bears (Ursus arctos) move in mountainous landscapes. We trained captive grizzly bears to walk on a horizontal treadmill and up and down 10% and 20% slopes. The cost of moving...
Carnahan, Anthony M.; van Manen, Frank T.; Haroldson, Mark A.; Stenhouse, Gordon; Robbins, Charles T. Thermal constraints on energy balance, behaviour and spatial distribution of grizzly bears
1. Heat dissipation limit theory posits that energy available for growth and reproduction in endotherms is limited by their ability to dissipate heat. In mammals, endogenous heat production increases markedly during gestation and lactation, and thus female mammals may be subject to greater thermal constraints on energy expenditure than males. Such...
Rogers, Savannah A.; Robbins, Charlie T.; Mathewson, Paul D.; Carnahan, Anthony M.; van Manen, Frank T.; Haroldson, Mark A.; Porter, Warren P.; Rogers, Taylor R.; Soule, Terrence; Long, Ryan A. Brown Bear (Ursus arctos; North America)
This chapter comprises the following sections: names, taxonomy, subspecies and distribution, descriptive notes, habitat, movements and home range, activity patterns, feeding ecology, reproduction and growth, behavior, parasites and diseases, status in the wild, and status in captivity.
Haroldson, Mark A.; Clapham, Melanie; Costello, Cecily M.; Gunther, Kerry A.; Kendall, Kate; Miller, Sterling; Pigeon, Karine; Proctor, Michael F.; Rode, Karyn D.; Servheen, Christopher; Stenhouse, Gordon; van Manen, Frank T. Whitebark pine cone production - 2020
No abstract available.
Haroldson, Mark A.; van Manen, Frank T. Yellowstone Grizzly Bear investigations 2019 - Annual report of the Interagency Grizzly Bear Study Team
No abstract available.
van Manen, Frank T.; Haroldson, Mark A.; Karabensh, Bryn Potential for recreational restrictions to reduce grizzly bear–caused human injuries
In 2011, 2 hikers were killed by grizzly bears (Ursus arctos) in separate incidents on backcountry trails in Hayden Valley, Yellowstone National Park, USA (YNP). Hayden Valley provides prime habitat for grizzly bears and is known to have high densities of bears. During 1970–2017, 23% (10 of 44) of all backcountry grizzly bear–inflicted human...
Gunther, Kerry A.; Haroldson, Mark A. Primarily resident grizzly bears respond to late-season elk harvest
Autumn ungulate hunting in the Greater Yellowstone Ecosystem carries the risk of hunter–grizzly bear (Ursus arctos) conflict and creates a substantial challenge for managers. For Grand Teton National Park, Wyoming, USA, a key information need is whether increased availability of elk (Cervus canadensis) carcasses during a late autumn (Nov–Dec)...
van Manen, Frank T.; Ebinger, Michael R.; Gustine, David D.; Haroldson, Mark A.; Wilmot, Katharine R.; Whitman, Craig Yellowstone Grizzly Bear investigations: Annual report of the Interagency Grizzly Bear Study Team 2018
This annual report summarizes the results of grizzly bear (Ursus arctos) research and monitoring conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2018. The research and monitoring program is focused on population estimation and demographics, food monitoring, and habitat monitoring. The...
van Manen, Frank T.; Haroldson, Mark A.; Karabensh, Bryn Estimating distemper virus dynamics among wolves and grizzly bears using serology and Bayesian state‐space models
Many parasites infect multiple hosts, but estimating the transmission across host species remains a key challenge in disease ecology. We investigated the within and across host species dynamics of canine distemper virus (CDV) in grizzly bears (Ursus arctos) and wolves (Canis lupus) of the Greater Yellowstone Ecosystem (GYE). We hypothesized that...
Cross, Paul C.; van Manen, Frank T.; Viana, Mafalda; Almberg, Emily S.; Bachen, Daniel; Brandell, Ellen E.; Haroldson, Mark A.; Hudson, Peter J.; Stahler, Daniel R.; Smith, Douglas W. Potential paths for male-mediated gene flow to and from an isolated grizzly bear population
For several decades, grizzly bear populations in the Greater Yellowstone Ecosystem (GYE) and the Northern Continental Divide Ecosystem (NCDE) have increased in numbers and range extent. The GYE population remains isolated and although effective population size has increased since the early 1980s, genetic connectivity between these populations...
Peck, Christopher P.; van Manen, Frank T.; Costello, Cecily M.; Haroldson, Mark A.; Landenburger, Lisa; Roberts, Lori L.; Bjornlie, Daniel D.; Mace, Richard D.