Jeffrey E Lovich, Ph.D.

As a Research Ecologist, Jeff is interested in the interactions of animals and their physical environment. Current research is focused on the ecology of turtles and tortoises and the effects of utility-scale renewable energy development (wind and solar) on wildlife, particularly in the Desert Southwest United States.

Biography

Jeff is a Research Ecologist with the Southwest Biological Science Center. He has studied the ecology and systematics of turtles and other reptiles for over 30 years, discovering and naming four of the world’s 356 turtle species, including three in the United States and one in Japan. Other interests include the ecological impacts of invasive species, the ecology and distribution of relict species, and the impacts of human activities (including renewable energy development) on ecological patterns and processes in the Mojave and Sonoran Deserts. During his career, he published numerous scientific papers and popular publications, including four books on topics ranging from turtles, to biological diversity, to the state of the Colorado River ecosystem in Grand Canyon. He serves on the editorial boards and directorates of various professional societies and he is a member of the IUCN (World Conservation Union), Species Survival Commission, Tortoise and Freshwater Turtle Specialist Group. He received a Fulbright Senior Specialist Award in 2008 to collaborate with researchers at Cadi Ayyad University in Marrakech, Morocco. He is an elected Fellow of The Linnean Society of London, the world's oldest active biological society. Current research focuses on the impacts of wind and solar energy development on federally protected populations of the desert tortoise and the ecology of poorly-known species of turtles in the United States and Morocco.

A native of Alexandria, Virginia, Jeff received his B.S. and M.S. degrees in Biology from George Mason University in 1982 and 1984, respectively, working with Carl Ernst. He received his Ph.D. in Ecology from the University of Georgia, Institute of Ecology in 1990 working with Whit Gibbons at the Savannah River Ecology Laboratory. Following graduate school, he worked for the Bureau of Land Management in southern California conducting research on the desert tortoise after their listing as a threatened species. He then became a charter member of the National Biological Survey, eventually merging with the U.S. Geological Survey, a better institutional home for an eclectic ecologist. After 15 years of conducting research on turtles and fish in rivers of the eastern United States, and another 10 years conducting research in the Mojave Desert of California, Jeff became a Research Manager, and subsequently, Center Director of the Western Ecological Research Center of the U.S. Geological Survey in Sacramento, California. He moved to Flagstaff in 2003 to become the Chief of the Grand Canyon Monitoring and Research Center of USGS. In 2005 he became the Deputy Director of the Southwest Biological Science Center in Flagstaff before returning to full time research in 2009. 

During his time as a practicing field researcher and manager Jeff published numerous scientific papers and popular publications. In addition he published four books. He is coauthor of "Turtles of the United States and Canada" published by the Smithsonian Institution Press in 1994; co-editor of, and contributor to, the book "Biological Diversity: Problems and Challenges," published by the Pennsylvania Academy of Science the same year; and co-editor and contributor to The State of the Colorado River Ecosystem in the Grand Canyon, published by the U.S. Geological Survey. His latest book, a substantially revised Turtles of the United States and Canada, Second Edition, was published by Johns Hopkins University Press in 2009. With 5,200 citations, it is the most comprehensive review of turtle biology ever written.

Jeff served on the editorial board or board of directors for several professional societies including the American Society of Ichthyologists and Herpetologists, the Herpetologists’ League, Herpetological Natural History, Herpetological Review, and the California Exotic Pest Plant Council. He remains on the editorial board of the journal Chelonian Conservation and Biology, and is a member of the IUCN (World Conservation Union), Species Survival Commission, Tortoise and Freshwater Turtle Specialist Group. He received a Fulbright Senior Specialist Award in 2008 to collaborate with researchers at Cadi Ayyad University in Marrakech, Morocco. He is an elected Fellow of The Linnean Society of London, the world's oldest active biological society. Jeff discovered, described and named four of the world’s 356 turtle species and subspecies, three of which occur in the United States. However science is just a hobby, he'd rather be spending time with his wife and kids, hunting, or driving one of his Pontiac GTO's.

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PUBLICATIONS

Lovich, J.E., and K.M. Hart. In press. Diamond-backed terrapin taxonomy: a history of controversy and uncertainty. In Roosenburg, W.M. and V.S. Kennedy (Eds.). Ecology and conservation of the diamondback terrapin Malaclemys terrapin. Johns Hopkins University Press, Baltimore, Maryland.

Lovich, J.E., J.W. Gibbons, and K. Green. In press. Life history of the diamond-backed terrapin with emphasis on geographic variation. In Roosenburg, W.M. and V.S. Kennedy (Eds.). Ecology and conservation of the diamondback terrapin Malaclemys terrapin. Johns Hopkins University Press, Baltimore, Maryland.

Lovich, J.E., J.R. Ennen, M. Agha, and J.W. Gibbons. 2018. Where have all the turtles gone, and why does it matter? BioScience. https://doi.org/10.1093/biosci/biy095

Lovich, J.E. 2018. Biographical sketch and bibliography of Jeffrey E. Lovich. Smithsonian Herpetological Information Service No. 153:1-31. https://repository.si.edu/handle/10088/35710

Lovich, J.E., M. Thomas, K. Ironside, C. Yackulic, and S.R. Puffer. 2018. Spatial distribution of estuarine diamond-backed terrapins (Malaclemys terrapin) and risk analysis from commercial blue crab (Callinectes sapidus) trapping at the Savannah Coastal Refuges Complex, USA. Ocean and Coastal Management 157:160-167. https://www.sciencedirect.com/science/article/pii/S0964569117308852

Lovich, J.E., S.R. Puffer, M. Agha, J.R. Ennen, K. Meyer-Wilkins, L.A. Tennant, A.L. Smith, T.R. Arundel, K.D. Brundige and M.S. Vamstad. 2018. Reproductive output and clutch phenology of female Agassiz's desert tortoises (Gopherus agassizii) in the Sonoran Desert region of Joshua Tree National Park. Current Herpetology 37(1):40–57.

Lovich, J.E., R.C. Averill-Murray, M. Agha, J.R. Ennen, and M. Austin. 2017. Variation in annual clutch phenology of Sonoran desert tortoises (Gopherus morafkai) in central Arizona. Herpetologica. http://www.hljournals.org/doi/10.1655/HERPETOLOGICA-D-17-00007.1

Zappalorti, R.T., A.M. Tutterow, S.E. Pittman, and J.E. Lovich. 2017. Hatching success and predation of bog turtle (Glyptemys muhlenbergii) eggs in New Jersey and Pennsylvania. Chelonian Conservation and Biology.

Ennen, J.R., W.A. Matamoros, M. Agha, J.E. Lovich, S.C. Sweat, and C.W. Hoagstrom. 2017. Hierarchical, quantitative biogeographic provinces for all North American turtles and their contribution to the biogeography of turtles and the continent. Herpetological Monographs 31:114-140. http://www.bioone.org/doi/10.1655/HERPMONOGRAPHS-D-16-00013

Moore-O’Leary, K.A., R.R. Hernandez, D.S. Johnston, S.R. Abella, K.E. Tanner, A.C. Swanson, J. Kreitler, and J.E. Lovich. 2017. Sustainability of utility-scale solar energy – critical ecological concepts. Frontiers in Ecology and the Environment. http://onlinelibrary.wiley.com/doi/10.1002/fee.1517/full

Lovich, J.E., and J.R. Ennen. 2017. Reptiles and amphibians. In, pp. 97-118. Perrow, M. (ed.), Wildlife and windfarms: conflicts and solutions. Vol. 1. Onshore. Pelagic Press. Exeter, U.K. https://pelagicpublishing.com/products/wildlife-and-wind-farms-vol-1-onshore-potential-effects

Agha, M., A.L. Smith, J.E. Lovich, D. Delaney, J.R. Ennen, J. Briggs, L.A. Tennant, S.R. Puffer, A. Walde, T.R. Arundel, S.J. Price, B.D. Todd. 2017. Mammalian mesocarnivore visitation at tortoise burrows in a wind farm. Journal of Wildlife Management. DOI: 10.1002/jwmg.21262. http://onlinelibrary.wiley.com/doi/10.1002/jwmg.21262/abstract

Lovich, J.E., M. Quillman, B. Zitt, A. Schroeder, D. Earl Green, C. Yackulic, P. Gibbons, and E. Goode. 2017. The effects of drought and fire in the extirpation of an abundant semi-aquatic turtle from a lacustrine environment in the southwestern USA. Knowledge and Management of Aquatic Ecosystems 418. http://www.kmae-journal.org/articles/kmae/abs/2017/01/kmae160179/kmae160179.html 

Ennen, J.R., J.E. Lovich, R.C. Averill-Murray, C. Yackulic, M. Agha, C. Loughran, L. Tennant, and B. Sinervo. 2017. The evolution of different maternal investment strategies in two closely related desert vertebrates. Ecology and Evolution. 1-13. http://onlinelibrary.wiley.com/doi/10.1002/ece3.2838/full 

Gibbons, J.W., J. Lovich and R.M. Bowden. 2017. Turtles: Freshwater, In Reference Module in Life Sciences, Elsevier, , ISBN: 978-0-12-809633-8, http://dx.doi.org/10.1016/B978-0-12-809633-8.01218-8 

Taniguchi, M., J.E. Lovich, K. Mine, S. Ueno, and N. Kamezaki. 2017. Unusual population attributes of invasive red-eared slider turtles (Trachemys scripta elegans) in Japan: do they have a performance advantage? Aquatic Invasions 12:97-108. http://www.aquaticinvasions.net/2017/AI_2017_Taniguchi_etal.pdf 

Lovich, J.E. and K. Yamamoto. 2016. Measuring the impact of invasive species on popular culture: a case study based on toy turtles from Japan. Humans and Nature (Museum of Nature and Human Activities, Hyogo, Japan) 27: 1-11. http://www.hitohaku.jp/publication/r-bulletin/2016-001.pdf 

Henderson, R.A., S.R. Puffer, J.E. Lovich. 2016. GOPHERUS AGASSIZII (Mohave Desert Tortoise). NEST DEPREDATION. Herpetological Review (Natural History Notes) 47(3):446-447.

Smith, A.L., S.R. Puffer, J.E. Lovich, L.A. Tennant, T.R. Arundel, M.S. Vamstad, and K.D. Brundige. 2016. A potential predator-prey interaction of an American badger and an Agassiz’s desert tortoise with a review of badger predation on turtles. California Fish and Game 102(3):131-144. https://nrm.dfg.ca.gov/FileHandler.ashx?DocumentID=136511&inline 

Lovich, J.E., M. Agha, C.W. Painter, L. Cole, A. Fitzgerald, K. Narum, and R. Jennings. 2016. Aspects of the reproductive ecology of female turtles in New Mexico. Western North American Naturalist 76. http://scholarsarchive.byu.edu/wnan/ 

Lovich, J.E. and J.R. Ennen. 2016. Energy development. In. pp. 31-34/ Jones, L.L.C, Halama, K.J., and Lovich, R.E. (eds.). Habitat management guidelines for amphibians and reptiles in the southwestern United States. Birmingham, Alabama, Partners in Amphibian and Reptile Conservation. Technical Publication HMG-5. http://www.swparc.org/2016/09/07/southwest-habitat-management-guideline-book-to-be-available-at-swparc-2016/

Lovich, J.E. 2016. Desert scrublands. In. Jones, L.L.C, Halama, K.J., and Lovich, R.E. (eds.). In, pp. 50-54. Habitat management guidelines for amphibians and reptiles in the southwestern United States. Birmingham, Alabama, Partners in Amphibian and Reptile Conservation. Technical Publication HMG-5. http://www.swparc.org/2016/09/07/southwest-habitat-management-guideline-book-to-be-available-at-swparc-2016/

Ernst, C.H., A. Laemmerzahl, and J.E. Lovich. 2016. A morphological review of subspecies of the Asian box turtle, Cuora amboinensis (Testudines, Geomydidae). Proceedings of the Biological Society of Washington 129:144-156. http://www.bioone.org/doi/10.2988/0006-324X-129.Q2.144

Ennen, J.R., J. Godwin, J.E. Lovich, B.R. Kreiser, B. Folt, and S. Hazard. 2016. Interdrainage morphological and genetic differentiation in the Escambia Map Turtle, Graptemys ernsti. Herpetological Conservation and Biology 11:122–131. [Link]

Ennen, J.R., M. Agha, W. Matamoros, S. Hazzard, and J.E. Lovich. 2016. Using climate, energy, and spatial-based hypotheses to interpret macroecological patterns of North America chelonians. Canadian Journal of Zoology 94:453–461. http://www.nrcresearchpress.com/doi/abs/10.1139/cjz-2016-0033#.VyEOWmfVyUk

Ennen, J.R., J.E. Lovich, and R.L. Jones. 2016. Graptemys pearlensis Ennen, Lovich, Kreiser, Selman, and Qualls 2010 – Pearl River Map Turtle. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs 5(9):094.1–8, doi:10.3854/crm.5.094.pearlensis.v1.2016, http://www.iucn-tftsg.org/cbftt/. [Download File]

Agha, M.A, M.O. Murphy, J.E. Lovich, J.R. Ennen, C.R. Oldham, K. Meyer, C. Bjurlin, M. Austin, S. Madrak, C. Loughran, L. Tennant, and S.J. Price. 2015. The effect of research activities and winter precipitation on defensive voiding behavior of Agassiz’s desert tortoises (Gopherus agassizii). Wildlife Research 41:641-649. CONTACT AUTHOR FOR A PDF

Lovich, J.E., J.R. Ennen, C.B. Yackulic, K. Meyer-Wilkins, M. Agha, C. Loughran, C. Bjurlin, M. Austin, and S. Madrak. 2015. Not putting all their eggs in one basket: bet-hedging despite extraordinary annual reproductive output of desert tortoises. Biological Journal of the Linnean Society 115:399–410. CONTACT AUTHOR FOR A PDF

Agha, M., B. Augustine, J.E. Lovich, D. Delaney, B. Sinervo, M.O. Murphy, J.R. Ennen, J.R. Briggs, R. Cooper, and S.J. Price. 2015. Using motion-sensor camera technology to infer seasonal activity and thermal niche of the desert tortoise (Gopherus agassizii). Journal of Thermal Biology 49-50:119-126. [Link]

Agha, M., J.E. Lovich, J.R. Ennen, B. Augustine, T.R. Arundel, M.O. Murphy, K. Meyer-Wilkins, C. Bjurlin, D. Delaney, J. Briggs, M. Austin, S. Madrak and S.J. Price. 2015. Turbines and terrestrial vertebrates: variation in tortoise survivorship between a wind energy facility and an adjacent undisturbed wildland area in the desert southwest (USA). Environmental Management. CONTACT AUTHOR FOR A PDF

Ennen, J.R., P.V. Lindeman, and J.E. Lovich. 2015. Intersexual allometry differences and ontogenetic shifts of coloration patterns in two aquatic turtles, Graptemys oculifera and Graptemys flavimaculata. Ecology and Evolution 5(11):2296–2305. [Link]

Kern, M.M., J.C. Guzy, J.E. Lovich, J.W. Gibbons, and M.E. Dorcas. 2015. Relationships of maternal body size and morphology with egg and clutch size in the diamondback terrapin, Malaclemys terrapin(Testudines: Emydidae). Biological Journal of the Linnean Society. CONTACT AUTHOR FOR A PDF

Lovich, J.E. 2015. Golden eagle mortality at a wind-energy facility near Palm Springs, California. Western Birds 46:76-80. [Download File]

Ernst, C.H. and J.E. Lovich. 2015. Interanal seam loss in Asian turtles of the Cuora flavomarginata complex (Testudines, Geoemydidae). Herpetological Bulletin 132:1-4. [Download File]

Lovich, J. 2015. Case study: road proliferation due to rapid renewable energy development. Chapter 4.5.1.1. In Andrews, K.A., P. Nanjappa, and S.P.D. Riley (Eds). Roads and Ecological Infrastructure: Concepts and Applications for Small Animals. Johns Hopkins University Press, Baltimore, Maryland. Pp. 79-80. [Download File]

Agha, M., D. Delaney, J.E. Lovich, J. Briggs, M. Austin and S.J. Price. 2015. Nelson’s big horn sheep (Ovis canadensis nelsoni) trample Agassiz’s desert tortoise (Gopherus agassizii) burrow at a California wind energy facility. Bulletin of the Southern California Academy of Sciences 114:58-62. [Download File]

Zappalorti, R.T., J.E. Lovich, R.F. Farrell, and M.E. Torocco. 2015. Nest-site characteristics of bog turtles (Glyptemys muhlenbergii) in New Jersey and Pennsylvania. Northeastern Naturalist 22:573–584.Any further reproduction or distribution of the manuscript, either in whole or in part, except for an individual’s own personal research purposes, is prohibited without written permission from the publisher, Eagle Hill Institute http://www.eaglehill.us/nena [Download File]

Lovich, J.E., J.W. Gibbons, and M. Agha. 2014. Does the timing of attainment of maturity influence sexual size dimorphism and adult sex ratio in turtles? Biological Journal of the Linnean Society 112:142-149. [Link]

Lovich, J.E., D. Delaney, J. Briggs, M. Agha, M. Austin, and J. Reese. 2014. Black bears (Ursus americanus) as a novel potential predator of Agassiz’s desert tortoises (Gopherus agassizii) at a California wind energy facility. Bulletin of the Southern California Academy of Sciences 113: 34–41. [Link]

Lovich, J. E. and J.R. Ennen. 2014. Graptemys gibbonsi Lovich and McCoy Pascagoula Map Turtle. Catalogue of American Amphibians and Reptiles. 901.1-901.8. [Link]

Godwin J., J.E. Lovich, J.R. Ennen, B.R. Kreiser, B.P. Folt, and C. Lechowicz. 2014. Hybridization of two megacephalic map turtles (Testudines: Emydidae: Graptemys) in the Choctawhatchee River drainage of Alabama and Florida. Copeia 2014:725-742. [Download File]

Ennen, J.R., M.E. Kalis, A.L. Patterson, B.R. Kreiser, J.E. Lovich, J. Godwin, and C.P. Qualls. 2014. Clinal variation or validation of a subspecies: a case study with the Graptemys nigrinoda complex (Testudines: Emydidae). Biological Journal of the Linnean Society. [Download File]

Lovich, J.E., Godwin, J.C., and McCoy, C.J. 2014. Graptemys pulchra Baur 1893 – Alabama Map Turtle. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5, pp. 072.1–072.6, doi:10.3854/crm.5.072.pulchra.v1.2014, http://www.iucn-tftsg.org/cbftt/. [Download File]

Lovich, J.E., C.T. LaRue, C.A. Drost, and T.R. Arundel. 2014. Traditional cultural use as a tool for inferring biogeography and provenance: a case study involving painted turtles (Chrysemys picta) and Hopi Native American Culture in Arizona, USA. Copeia 2014:215-220. [Download File]

Lovich, J.E., C.H. Ernst, E.M. Ernst and J.L. Riley. 2014. A 21 year study of seasonal and interspecific variation of hatchling emergence in a Nearctic freshwater turtle community: to overwinter or not to overwinter. Herpetological Monographs 28:93-109. [Download File]

Lovich, J.E., M. Agha, C.B. Yackulic, K. Meyer, C. Bjurlin, J.R. Ennen, T.R. Arundel, and M. Austin. 2014. Nest site characteristics, nesting movements, and lack of long-term nest site fidelity in Agassiz’s desert tortoises (Gopherus agassizii) at a wind energy facility in southern California. California Fish and Game 100:404-416. [Download File]

Lovich, J.E., C.B. Yackulic, J. Freilich, M. Agha, M. Meulblok, K.P. Meyer, T.R. Arundel, J. Hansen, M.S. Vamstad and S.A. Root. 2014. Climatic variation and tortoise survival: has a desert species met its match? Biological Conservation 169:214-224. [Link]

Lovich, J.E. 2013(2014). Assessing the long-term survival and reproductive output of desert tortoises at a wind energy facility near Palm Springs, California - Final Project Report. California Energy Commission. Publication number: CEC-500-2014-005. 60 pp. [Link]

Lovich, J.E. and J.R. Ennen. 2013. Assessing the state of knowledge of utility-scale wind energy development and operation on non-volant terrestrial and marine wildlife. Applied Energy 103:52–60.[Download File]

Lovich, J.E. and J.R. Ennen. 2013. A quantitative analysis of the state of knowledge of turtles of the United States and Canada. Amphibia-Reptilia. 34:11-23. [Download File]

Agha, M., Lovich, J.E., Ennen, J.R., and Wilcox, E. 2013. Nest-guarding by female Agassiz's desert tortoise (Gopherus agassizii) at a wind-energy facility near Palm Springs, California. Southwestern Naturalist 58(2): 254-257. [Download File]

Underwood, E.B., S. Bowers, J.C. Guzy, J.E. Lovich, C.A. Taylor, J.W. Gibbons, and M.E. Dorcas. 2013. Sexual dimorphism and feeding ecology of diamondback terrapins (Malaclemys terrapin). Herpetologica 69:397-404. [Download File]

Lovich, J.E., N.J. Scott, R.B. Bury, K.C. Dodd, Jr., and R.W. McDiarmid. 2012. A history of herpetologists and herpetology in the U.S. Department of the Interior. Herpetological Conservation and Biology 7(Monograph 2):1-45. [Link]

Lovich, J.E. 2012. Assessing desert tortoise survival and reproduction at a wind energy facility near Palm Springs, California. California Energy Commission, Public Interest Energy Research (PIER) Program, Environmental Area Research. [Link]

Ennen, J.R., J.E. Lovich, K. Meyer, C. Bjurlin, and T.R. Arundel. 2012. Nesting ecology of a Desert Tortoise (Gopherus agassizii) population at a utility-scale renewable energy facility in southern California. Copeia 2012:222-228. [Download File]

Ennen, J.R., K.P. Meyer, and J.E. Lovich. 2012. Female Agassiz’s desert tortoise activity at a wind energy facility in southern California: the influence of an El Niño event. Natural Science 4:30-37. doi:10.4236/ns.2012.41006 [Download File]

Naimi, M., M. Znari, J.E. Lovich, Y. Feddadi, M. A. Ait Baamrane. 2012. Clutch and egg allometry of the turtle Mauremys leprosa (Chelonia: Geoemydidae) from a polluted peri-urban river in west-central Morocco. Herpetological Journal 22:43-49. [Download File]

Lovich, J.E., S.V. Madrak, C. Drost, A.J. Monatesti, D. Casper, and M. Znari. 2012. Optimal egg size in a suboptimal environment: reproductive ecology of female Sonora mud turtles (Kinosternon sonoriense) in central Arizona, USA. Amphibia-Reptilia 33:161-170. [Download File]

Lovich, J.E., M. Agha, M. Meulblok, K. Meyer, J. Ennen, C. Loughran, S.V. Madrak, and C. Bjurlin. 2012. Climatic variation affects clutch phenology in Agassiz's Desert Tortoise (Gopherus agassizii). Endangered Species Research 19:63–74. [Download File]

Ennen, J.R., J.E. Lovich and R.Birkhead. 2012. Graptemys pearlensis. Ennen, Lovich, Kreiser, Selman, and Qualls. Pearl River Map Turtle. Catalogue of American Amphibians and Reptiles. 889.1-889.4.[Download File]

Lovich, J.E., J.R. Ennen, S. Madrak, K. Meyer, C. Loughran, C. Bjurlin, T. Arundel, W. Turner, C. Jones, and G.M. Groenendaal. 2011. Effects of wind energy production on growth, demography and survivorship of a desert tortoise (Gopherus agassizii) population in southern California with comparisons to natural populations. Herpetological Conservation and Biology 6:161-174. [Link]

Drost, C.A, J.E. Lovich, S.V. Madrak and A.J. Monatesti. 2011. Removal of nonnative slider turtles (Trachemys scripta) and effects on native Sonora mud turtles (Kinosternon sonoriense) at Montezuma Well, Yavapai County, Arizona. U.S. Geological Survey Open-File Report 2010–1177, 48 p. [Link]

Lovich, J.E., J.R. Ennen, S. Madrak, C. Loughran, K. Meyer, T. Arundel, and C. Bjurlin. 2011. Long-term post fire effects on spatial ecology and reproductive output of female desert tortoises at a wind energy facility near Palm Springs, California. Fire Ecology 7:75-87. [Download File]

Lovich, J.E. and J.R. Ennen. 2011. Wildlife conservation and solar energy development in the Desert Southwest, United States. BioScience 61:982-992. [Download File]

Lovich, J.E. and G.B. Haxel. 2011. A previously unreported locality record for the Gila monster. Bulletin of the Southern California Academy of Sciences 110:59-62. [Download File]

Ernst, C.H., A.F. Laemmerzahl, and J.E. Lovich. 2011. Does the "kamaroma"-plastron pattern morph occur in both Philippine subspecies of the turtle Cuora amboinensis? Proceedings of the Biological Society of Washington 124:259-269.011 [Download File]

Lovich, J.E., J.R. Ennen, S.V. Madrak, and B. Grover. 2011. Turtles, culverts and alternative energy development: an unreported but potentially significant mortality threat to the desert tortoise (Gopherus agassizii). Chelonian Conservation and Biology 10:124-129. [Download File]

Ennen, J.R., R.D. Birkhead, B.R. Kreiser, D.L. Gaillard, C.P. Qualls, and J.E. Lovich. 2011. The effects of isolation on the demography and genetic diversity of long-lived species: implications for conservation and management of the gopher tortoise (Gopherus polyphemus). Herpetological Conservation and Biology 6:202-214. [Download File]

Lovich, J.E., Yasukawa, Y., and Ota, H. 2011. Mauremys reevesii (Gray 1831) – Reeves’ Turtle, Chinese Three-keeled Pond Turtle. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5, pp. 050.1–050.10, doi:10.3854/crm.5.050.reevesii.v1.2011 [Download File]

Lovich, J.E., Godwin, J.C., and McCoy, C.J. 2011. Graptemys ernsti Lovich and McCoy 1992 – Escambia Map Turtle. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5, pp. 051.1–051.6, doi:10.3854/crm.5.051.ernsti.v1.2011 [Download File]

Ennen, J.R., C.L. Loughran, and J.E. LovichGopherus agassizii (Desert Tortoise). Non-native seed dispersal. Herpetological Review (Natural History Notes) 42:266-267. [Download File]

Lovich, J.E. 2011. Gopherus agassizii (Desert Tortoise) and Crotalus ruber (Red Diamond Rattlesnake) burrow co-occupancy. Herpetological Review. 42:421. [Download File]

Loughran, C.L., J.E. Ennen, and J.E. Lovich. 2011. Gopherus agassizii (Desert tortoise). Burrow collapse. Herpetological Review 42:593 [Download File]

Lovich, J.E., C. Drost, A.J. Monatesti, D. Casper, D. Wood, and M. Girard. 2010. New reptilian prey items for the Sonora mud turtle (Kinosternon sonoriense) with a brief review of saurophagy and ophiophagy in North American turtles. Southwestern Naturalist 55:135-138. [Download File]

Ennen, J.R.,J.E. Lovich, B.R. Kreiser, W. Selman, and C.P. Qualls. 2010. Genetic and morphological variation between populations of the Pascagoula Map Turtle (Graptemys gibbonsi) in the Pearl and Pascagoula Rivers with description of a new species. Chelonian Conservation and Biology 9:98-113. [Download File]

Ennen, J.R.,B.R. Kreiser, C.P. Qualls, and J.E. Lovich. 2010. A morphological and molecular reassessment of Graptemys oculifera and G. flavimaculata. Journal of Herpetology 44:544-554. [Download File]

Lovich, J.E., M. Znari, M. Abdeljalil Ait Baamrane, M. Naimi, A. Mostalih. 2010. Biphasic geographic variation in sexual size dimorphism of turtle (Mauremys leprosa) populations along an environmental gradient in Morocco. Chelonian Conservation and Biology 9:45-53. [Download File]

Ernst, C.H. and J.E. Lovich. 2009. Turtles of the United States and Canada. Second edition. Johns Hopkins University Press. 827 pages. [Link]

Glenn, E.P, P.L. Nagler and J.E. Lovich. 2009. The surprising value of saltcedar. Southwest Hydrology May/June issue: 10-11. [Download File]

Lovich, J.E., W. Selman, and C.J. McCoy. 2009. Graptemys gibbonsi Lovich and McCoy 1992 – Pascagoula map turtle, Pearl River map turtle, Gibbon’s map turtle. In, Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.). Conservation biology of freshwater turtles and tortoises: a compilation project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5, pp. 029.1-029.8, doi:10.3854/crm.5.029.gibbonsi.v1.2009. [Download File]

Ernst, C.H., A.E. Laemmerzahl and J.E. Lovich. 2008. A morphological review of the Cuora flavomarginata complex (Testudines: Geoemydidae). Proc. Biol. Soc. 121(3):391-397. [Download File]

Lovich, J.E. and K.R. Beaman. 2008. Distribution of native turtles in the arid southwestern United States with comments on Kinosternon sonoriense: a species presumed to be lost from California’s herpetofauna. In, pp. 127-134, Reynolds, R.E. (ed.). The 2008 Desert Symposium and Field Guide and Proceedings. California State University, Desert Studies Consortium and LSA Associates, Inc.[Download File]

Lovich, J.E. and K.R. Beaman. 2007. A history of Gila monster (Heloderma suspectum cinctum) records from California with comments on factors affecting their distribution. Bulletin of the Southern California Academy of Science. 106(2):39-58. [Download File]

Lovich, J.E. and T. S. Melis. 2007. The state of the Colorado River ecosystem in Grand Canyon: lessons from 10 years of adaptive ecosystem management. International Journal of River Basin Management. 5(3):207-221. [Download File]

Gloss, S.P., J.E. Lovich, and T.S. Melis (eds.). 2005. The state of the Colorado River ecosystem in the Grand Canyon. U.S. Geological Survey Circular 1282. 220 pages. http://pubs.usgs.gov/circ/1282/.[Link]

Lovich, J.E. and T.S. Melis. 2005. Lessons from 10 years of adaptive management in Grand Canyon. In, pp. 207-220, Gloss, S.P., Lovich, J.E., and Melis, T.S., (eds.). The state of the Colorado River ecosystem in the Grand Canyon: U.S. Geological Survey Circular 1282. [Download File]

Lovich, J.E. Profiles of selected fish species found in the Grand Canyon ecosystem. In, pp. 50-53, Gloss, S.P., Lovich, J.E., and Melis, T.S., (eds.). The state of the Colorado River ecosystem in the Grand Canyon: U.S. Geological Survey Circular 1282. 220 pages. [Download File]

Lovich, J.E. Diamondback Terrapin Still Vulnerable to Human Activities. Sound Waves (monthly newsletter produced by the U.S. Geological Survey, online from January 1999 to present. December 2002/January 2003 issue. pp. 7-9. [Link]

Lovich, J. and K. Meyer. 2002. The western pond turtle (Clemmys marmorata) in the Mojave River, California, USA: Highly adapted survivor or tenuous relict? Journal of Zoology, London 256:537-545.[Download File]

Gibbons, J. W., J.E. Lovich, A. D. Tucker, N. N. FitzSimmons, and J. L. Greene. 2001. Demographic and ecological factors affecting conservation and management of the diamondback terrapin (Malaclemys terrapin) in South Carolina. Chelonian Conservation and Biology 4(1):66-74. [Download File]

Lovich, J. E. and R. Daniels. 2000. Environmental characteristics of desert tortoise (Gopherus agassizii) burrow locations in an altered industrial landscape. Chelonian Conservation and Biology 3(4):714-721. [Download File]

Nicolai, N. C. and J. E. Lovich. 2000. Preliminary observations of the behavior of male, flat-tailed horned lizards before and after an off-highway vehicle race in California. California Fish and Game 86(3):208-212. [Download File]

Burke, V. J., J. E. Lovich, and J. W. Gibbons. 2000. Conservation of freshwater turtles. In, pp. 156-179, M. Klemens (ed.), Turtle Conservation. Smithsonian Institution Press, Washington, D.C. [Download File]

Dudley, T.L., C.J. DeLoach, J.E. Lovich, and R.I. Carruthers. 2000. Saltcedar invasion of western riparian areas: impacts and new prospects for control. In, pp. 345-381, E. McCabe and S. E. Loos (eds.), Trans. 65th No. Amer. Wildl. and Natur. Resour. Conf. Wildlife Management Institute, Washington, D.C. [Download File]

DeLoach, C.J., R.I. Carruthers, J.E. Lovich, T.L. Dudley, and S.D. Smith. 2000. Ecological interactions in the biological control of saltcedar (Tamarix spp.) in the United States: toward a new understanding.In, pp. 819-873, N. R. Spencer (ed.), Proceedings of the X International Symposium on Biological Control of Weeds. 4-14 July, 1999, Montana State University, Bozeman, Montana. [Download File]

Lovich, J. E. and D. Bainbridge. 1999. Anthropogenic degradation of the southern California desert ecosystem and prospects for natural recovery and restoration. Environmental Management 24:309-326.[Download File]

Lovich, J. E., P. Medica, H. Avery, K. Meyer, G. Bowser, and A. Brown. Studies of reproductive output of the desert tortoise at Joshua Tree National Park, the Mojave National Preserve, and comparative sites. Park Science 19:22-24. (Reprinted in Tortuga Gazette [Newsletter of the California Turtle and Tortoise Club] 1999. 35(7):1-2, 4. [Download File]

Lovich, J. E., C. H. Ernst, R. T. Zappalorti, and D. W. Herman. 1998. Geographic variation in growth and sexual size dimorphism of bog turtles (Clemmys muhlenbergii). American Midland Naturalist 139:69-78. [Download File]

Lovich, J. E. and R. G. de Gouvenain. 1998. Saltcedar invasion in desert wetlands of the southwestern United States: Ecological and political implications. In, pp. 447-467, S. K. Majumdar, E. W. Miller, and F. J. Brenner (eds.), Ecology of Wetlands and Associated Systems. Pennsylvania Academy of Science. [Download File]

Gibbons, J. W., V. J. Burke, J. E. Lovich, R. D. Semlitsch, T. D. Tuberville, J. R. Bodie, J. L. Greene, P. H. Niewiarowski, H. H. Whiteman, D. E. Scott, J. H. K. Pechmann, C. R. Harrison, S. H. Bennett, J. D. Krenz, M. S. Mills, K. A. Buhlmann, J. R. Lee, R. A. Seigel, A. D. Tucker, T. M. Mills, T. Lamb, M. E. Dorcas, J. D. Congdon, M. H. Smith, D. H. Nelson, M. B. Deitsch, H. H. Hanlin, J. A. Ott, and D. J. Karapatakis. 1997. Perceptions of species abundance, distribution, and diversity: lessons from four decades of sampling on a government-managed reserve. Environmental Management 21:259-268.[Download File]

Lovich, J. E. 1997. Additional records of amphibians and reptiles from Indiana County, Pennsylvania. J. Pennsylvania Acad. Sci. 71:35-38. [Download File]

Lovich, J. E. and J. W. Gibbons. 1997. Conservation of covert species: protecting species we don't even know. In, pp. 426-429, J. Van Abbema et al. (eds.), Proceedings: Conservation, Restoration and Management of Turtles and Tortoises - an International Conference, State University of New York, Purchase. [Download File]

Ernst, C. H., J. E. Lovich, A. F. Laemmerzahl, and S. Sekscienski. 1997. A comparison of plastron scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian Conservation and Biology 2:603-607. [Download File]

Hinton, T. G., P. Fledderman, J. Lovich, J. Congdon, and J. W. Gibbons. 1997. Radiographic determination of fecundity: Is the technique safe for developing turtle embryos? Chelonian Conservation and Biology 2:409-414. [Download File]

Lovich, J. E. Wildlife as weeds. 1997. In, pp. 46-51, J. Lovich, J. Randall, and M. Kelly (eds.), Proceedings California Exotic Pest Plant Council Symposium '96. [Download File]

Lovich, J. E., S. W. Gotte, C. H. Ernst, J. Harshbarger, A. F. Laemmerzahl, and J. W. Gibbons. 1996. Prevalence and histopathology of shell disease in turtles from Lake Blackshear, Georgia. J. Wildl. Diseases 32:259-265. [Download File]

Lovich, J. E. and R. E. Lovich. 1996. Decline of native brook trout (Salvelinus fontinalis) populations along the upper West Branch of the Susquehanna River: Canaries outside the coal mine. J. Pennsylvania Acad. Sci. 70:55-60. [Download File]

Lovich, J. E. 1996. Possible demographic and ecologic consequences of sex ratio manipulation in turtles. Chelonian Conservation and Biology 2:114-117. [Download File]

Sloan, K. N., K. A. Buhlmann, and J. E. Lovich. 1996. Stomach contents of commercially harvested adult alligator snapping turtles, Macroclemys temminckii. Chelonian Conservation and Biology 2:96-99.[Download File]

Lovich, J. E. and T. Lamb. 1995. Morphometric similarity between the turtles Kinosternon subrubrum hippocrepis and K. baurii. J. Herpetology 29:621-624. [Download File]

Lovich, J. E. Ecology of North American tortoises. 1995. In, G. Aguirre, E. D. McCoy, H. Mushinsky, M. Villagrán Santa Cruz, R. García, and G. C. Andreu (eds.), Proceedings of North American Tortoise Conference, Mapimi Biosphere Reserve, Durango, Mexico, October 8-11, 1994. Publications de la Sociedad Herpetologica Mexicana No. 2:50-53. [Download File]

Sloan, K. and J. E. Lovich. 1995. Exploitation of the alligator snapping turtle, Macroclemys temminckii, in Louisiana: A case study. Chelonian Conservation and Biology 1:221-222. [Download File]

Lovich, J. E. and C. J. McCoy. 1994. Graptemys ernsti. Cat. Amer. Amphib. Rept. 585.1-585.2 [Download File]

Lovich, J. E. and C. J. McCoy. 1994. Graptemys gibbonsi. Cat. Amer. Amphib. Rept. 586.1-586.2 [Download File]

Lovich, J. E. Biodiversity and zoogeography of non-marine turtles in Southeast Asia. 1994. In, pp. 380-391, Biological Diversity: Problems and Challenges. S. K. Majumdar, F. J. Brenner, J. E. Lovich, E. W. Miller, and J. F. Schalles (eds.), Pennsylvania Academy of Science. [Download File]

Lovich, J. E. 1993. Macroclemys temminckii. Cat. Amer. Amphib. Rept. 562.1-562.4. [Download File]

Lovich, J. E. and C. J. McCoy. 1992. Review of the Graptemys pulchra group (Reptilia, Testudines, Emydidae), with descriptions of two new species. Annals of Carnegie Museum 61:293-315. [Download File]

Lovich, J. E., D. W. Herman, and K. M. Fahey. 1992. Seasonal activity and movements of bog turtles (Clemmys muhlenbergii) in North Carolina. Copeia 1992:1107-1111. [Download File]

Grant, B.W., A.D. Tucker, J.E. Lovich, A.M. Mills, P.M. Dixon, and J.W. Gibbons. 1992. The use of coverboards in estimating patterns of reptile and amphibian biodiversity. Pp. 379-403 in Wildlife 2001: Populations. D. L. McCullough and R. H. Barrett (eds.), Elsevier Applied Science, London. [Download File]

Ross, D.A. and J.E. Lovich. 1992. Does the color pattern of two species of turtles imitate duckweed? J. Pennsylvania Acad. Sci. 66:39-42. [Download File]

Lovich, J.E. 1992. Aspects of the ecology of an isolated brook trout (Salvelinus fontinalis) population in Fairfax County, Virginia. J. Pennsylvania Acad. Sci. 65:107-111. [Download File]

Lovich, J. E. and J. W. Gibbons. 1992. A review of techniques for quantifying sexual size dimorphism. Growth, Development and Aging 56:269-281. [Download File]

Lovich, J.E., A.F. Laemmerzahl, C.H. Ernst, and J.F. McBreen. 1991. Relationships among turtles of the genus Clemmys (Reptilia:Testudines:Emydidae) as suggested by plastron scute morphology. Zoologica Scripta 20:425-429. [Download File]

Lovich, J.E., A.D. Tucker, D.E. Kling, J.W. Gibbons, and T.D. Zimmerman. 1991. Behavior of hatchling diamondback terrapins released in a South Carolina salt marsh. Herpetological Review 22:81-83.[Download File]

Garstka, W.R., W.E. Cooper, Jr., K.W. Wasmund, and J.E. Lovich. 1991. Males sex steroids and hormonal control of male courtship behavior in the yellow‑bellied slider turtle, Trachemys scripta. Comp. Biochem. Physiol. 98A: 271‑280. [Download File]

Gibbons, J.W. and J.E. Lovich. 1990. Sexual dimorphism in turtles with emphasis on the slider turtle (Trachemys scripta). Herpetological Monographs 4:1‑29. [Download File]

Lamb, T. and J.E. Lovich. 1990. Morphometric validation of the striped mud turtle (Kinosternon baurii) in the Carolinas and Virginia. Copeia 1990:613‑618. [Download File]

Lovich, J.E., C.J. McCoy and W.R. Garstka. 1990. The development and significance of melanism in the slider turtle. Pp. 233-254 in Life History and Ecology of the Slider Turtle. J. Whitfield Gibbons (ed.), Smithsonian Institution Press, Washington, DC. [Download File]

Lovich, J., J.W. Garstka, and W. Cooper. 1990. Female participation in courtship behavior of the turtle, Trachemys scripta scripta. J. Herpetology 24:422‑424. [Download File]

Lovich, J.E. 1990. Spring movement patterns of two radio‑tagged male spotted turtles. Brimleyana 16:67‑71. [Download File]

Lovich, J. E. and J. W. Gibbons. 1990. Age at maturity influences adult sex ratio in the turtle Malaclemys terrapin. Oikos 59:126‑134. [Download File]

Ernst, C.H. and J.E. Lovich. 1990. A new species of Cuora (Reptilia: Testudines: Emydidae) from the Ryukyu Islands. Proc. Biol. Soc. Washington 103: 26‑34. [Download File]

Lovich, J.E. 1990. Gaping behavior in basking eastern painted turtles. Journal of the Pennsylvania Academy of Science 64:78‑80. [Download File]

Lovich, J.E., C.H. Ernst, and J. McBreen. 1990. Growth, maturity and sexual dimorphism in the wood turtle. Canadian J. Zool. 68:672‑677. [Download File]

Lovich, J.E., and C.H. Ernst. 1989. Variation in the plastral formulae of selected turtles with comments on taxonomic utility. Copeia 1989:304‑318. [Download File]

Iverson, J.B., C.H. Ernst, S.W. Gotte, and J.E. Lovich. 1989. The validity of Chinemys megalocephala (Testudines: Batagurinae). Copeia 1989:494‑498. [Download File]

Ernst, C.H., R.T. Zappalorti, and J.E. Lovich. 1989. Overwintering sites and thermal relations of hibernating bog turtles, Clemmys muhlenbergii. Copeia 1989:761‑764. [Download File]

Lovich, J.E. 1989. The spotted turtles of Cedar Bog, Ohio: historical analysis of a declining population. Pp. 23-28 in Proceedings of Cedar Bog Symposium II. R.C. Glotzhober, A. Kochman, and W.T. Schultz (eds.), Ohio Historical Society. [Download File]

Lovich, J.E. 1989. The Savannah River Ecology Laboratory Herpetological Museum. Herpetological Review 20:37-38. [Download File]

Lovich, J.E. 1989. Another exotic turtle record for Hawaii? 'Elepaio (J. Hawaii Audubon Soc.). 49: 86-87. [Download File]

Lovich, J.E. 1988. Geographic variation in the seasonal activity cycle of spotted turtles, Clemmys guttata. J. Herpetol. 22:482‑485. [Download File]

Lovich, J.E. 1988. Aggressive basking behavior in eastern painted turtles (Chrysemys picta picta). Herpetologica 44:197‑202. [Download File]

Lovich, J.E., and T. R. Jaworski. 1988. Annotated checklist of amphibians and reptiles reported from Cedar Bog, Ohio. Ohio J. Sci. 88: 139-143. [Download File]

Sanderson, R.A. and J.E. Lovich. 1988. Graptemys barbouri. Cat. Amer. Amphib. Rept. 421. 1-421. 2. [Download File]

Lovich, J.E. 1987. Mountain nightingales: The story of wolves in western Pennsylvania. Mountain Journal 5:3-7. [Download File]

Lovich, J.E. 1986. Daily activity cycles of introduced Hawaiin birds at an urban roost. 'Elepaio (J. Hawaii Aubudon Soc.) 46: 131-132. [Download File]

Ernst, C.H., and J.E. Lovich. 1986. Morphometry in the chelid turtle, Platemys platycephala. Herpetological Journal 1: 66-70. [Download File]

Ernst, C.H., S.W. Gotte, and J.E. Lovich.1985. Reproduction in the mole kingsnake, Lampropeltis calligaster calligaster. Bull. Maryland Herpetol. Soc. 21: 16‑22. [Download File]

Lovich, J.E., C.H. Ernst, and S.W. Gotte. 1985. Geographic variation in Chinemys reevesii (Gray) and the status of Geoclemys grangeri Schmidt. J. Herpetol. 19:238‑245. [Download File]

Lovich, J.E. Graptemys pulchra. 1985. Cat. Amer. Amphib. Rept. 360.1‑360.2. [Download File]