Dr. Sasha Reed is an ecologist focused on understanding how our planet's ecosystems work and what factors determine the services they provide. The study sites and methods Sasha uses are diverse, and with each of her projects she strives to provide scientific information that helps our nation address challenges, solve problems, and maximize opportunities.
Sasha seeks to provide useful information for land managers, policy-makers, and the general public. Sasha uses both basic and applied scientific approaches to improve our understanding of the fundamental controls over ecosystems, to determine how these systems respond to change, and to explore solutions for addressing relevant problems. Sasha works closely with a range of collaborators - including federal agency partners (BLM, NPS, DOE, DoD, BIA, USFS, USFWS) - in designing research studies, conducting information and technology transfer, and performing outreach activities. Some of her primary research interests include understanding how drought and increasing temperatures affect ecosystems, exploring a diversity of energy options for meeting national demand, assessing the consequences of exotic plant invasion and ways to combat them, and establishing novel management options for increased effectiveness and efficiency in restoration and reclamation. Sasha attempts to conduct research that is innovative, collaborative, and useful.
Professional Experience
Research Ecologist, USGS, Southwest Biological Science Center, Moab, UT: May 2008 - present
Research Ecologist, USGS-SCEP Program, Moab, UT: 2005 - 2007 summers only
Fellow, NSF Graduate Research Fellowship: Awarded April 2003; deferred until September 2005-May 2008
Fellow, National Science Foundation IGERT Fellowship, Carbon Climate and Society Initiative (CCSI): August 2003-August 2005.
Education and Certifications
Ph.D. in Biogeochemistry, 2002-2008: University of Colorado at Boulder, Department of Ecology & Evolutionary Biology and Institute of Arctic & Alpine Research (INSTAAR).
B.A. in Organic Chemistry, 1993-1997: Colgate University, Department of Chemistry, Hamilton, NY. Graduated magna cum laude.
Honors and Awards
Elected Member-At-Large, Ecological Society of America (ESA). August 2020-August 2022.
Ecological Society of America (ESA) Early Career Fellow. March 2016.
Presidential Early Career Award for Scientists and Engineers (PECASE). October 2011.
Star Award, Department of the Interior. October 2010, May 2000.
Graduate Student Research and Creative Works Award, University of Colorado at Boulder. May 2008.
USGS Student Career Experience Program (SCEP) Fellowship. June-August of 2006 & 2007.
Student Policy Award, Ecological Society of America (ESA). September 2006.
AAAS Program for Excellence in Science. August 2006.
Lawrence Award. April 1996.
Phi Ea Sigma University Honor Society Awarded Membership. April 1994-May 1997.
Phi Eta Sigma Aid Committee Selected Board Member. September 1995-May 1997.
Barry Goldwater Award Nominee. November 1994.
Science and Products
Regional Assessment of Drought Impacts on Soils (RADIS)
Climate Adaptation Strategies for Arid Grasslands
Biological Soil Crust ("Biocrust") Science
Diverse data to improve Southwest fire forecasts: Joining novel remote sensing, post-fire dynamics, and intra-annual precipitation patterns
Using a multi-scale approach to synthesize measurements and models of C4 photosynthesis
GrassCast: A multi-agency tool using remote sensing, modeling, and on-the-ground science to forecast grassland productivity in the Southwest
Learning From the Past and Planning for the Future: Experience-Driven Insight Into Managing for Ecosystem Transformations Induced by Drought and Wildfire
Determining Successful Management and Restoration Strategies for Pinyon-Juniper Communities in the Face of Changing Climate and Wildfire
Implementing a Grassland Productivity Forecast Tool for the U.S. Southwest
Long-Term Vegetation Change on the Colorado Plateau
Novel multi‐scale synthesis of nitrogen fixation rates and drivers across the terrestrial biosphere
Southwest Energy Exploration, Development, and Reclamation (SWEDR)
Meteorological measurements from five weather stations in Grand and San Juan Counties in southeastern Utah (ver. 2.0, April 2023)
Ecological property data and experimental lab incubation results from a long-term nitrogen deposition simulation experiment in three semi-arid grasslands, Arches National Park, Utah, USA, 2013-2019
Data and software code from two long-term experiments (1996-2011 and 2005-2018) at three sites on the Colorado Plateau of North America
CO2 concentrations and microbial biomass data derived from incubation experiments on soils collected at Arches National Park in 2017 and 2018
Riparian vegetation, topography, and ground cover constituents along the Upper Colorado River near Moab, UT (2010-2017) (ver. 1.1, Jan 2023)
Erosion and Rehabilitation Data, Bandelier National Monument, New Mexico, USA
Time, climate, and soil settings set the course for reclamation outcomes following dryland energy development
Storms and pH of dam releases affect downstream phosphorus cycling in an arid regulated river
Community for data integration 2019 project report
Long-term relationships between seed bank communities and wildfire across four North American desert sites
Droughting a megadrought: Ecological consequences of a decade of experimental drought atop aridification on the Colorado Plateau
Soil surface treatments and precipitation timing determine seedling development across southwestern US restoration sites
Grazing and ecosystem service delivery in global drylands
Exploring the role of cryptic nitrogen fixers in terrestrial ecosystems: A frontier in nitrogen cycling research
Patterns and controls of foliar nutrient stoichiometry and flexibility across United States forests
Understory plant communities show resistance to drought, hurricanes, and experimental warming in a wet tropical forest
Southeast Utah Group climate and drought adaptation report: Exposure and perennial grass sensitivity
What is a biocrust? A refined, contemporary definition for a broadening research community
Non-USGS Publications**
**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
Science and Products
- Science
Filter Total Items: 17
Regional Assessment of Drought Impacts on Soils (RADIS)
Soils are the foundation of terrestrial ecosystems. They provide critical services including supplying a substrate and the nutrients necessary for plant growth, retaining moisture from precipitation, filtering contaminants from percolating waters, and acting as a sink of carbon. Healthy soils are key to sustaining both human and ecosystem health. However, global- and regional-scale disturbances...Climate Adaptation Strategies for Arid Grasslands
Helping National Parks in the 4-Corners region manage grasslands in a changing climate.Biological Soil Crust ("Biocrust") Science
Biological soil crusts (biocrusts) are commonly found on the soil surface in arid and semi-arid ecosystems (collectively called drylands). Biocrusts can consist of mosses, cyanobacteria, lichens, algae, and microfungi, and they strongly interact with the soil. These organisms or consortium of disparate organisms, depending on the specific biocrust, are important to the functioning of ecosystems...Diverse data to improve Southwest fire forecasts: Joining novel remote sensing, post-fire dynamics, and intra-annual precipitation patterns
Fire has increased dramatically across the western U.S. and these increases are expected to continue. With this reality, it is critical that we improve our ability to forecast the timing, extent, and intensity of fire to provide resource managers and policy makers the information needed for effective decisions. For example, an advanced, spatially-explicit prediction of the upcoming fire season wouUsing a multi-scale approach to synthesize measurements and models of C4 photosynthesis
Plants convert carbon dioxide into sugars for food during photosynthesis, and this provides food for all animal life. However, photosynthesis is inhibited when a plant’s enzymes use oxygen instead of carbon dioxide. To avoid this use of oxygen, some plants developed a photosynthetic adaptation – called C4 photosynthesis – to concentrate carbon dioxide around the enzymes. While less than 5% of plan...GrassCast: A multi-agency tool using remote sensing, modeling, and on-the-ground science to forecast grassland productivity in the Southwest
Rangeland ecosystems are one of the largest single providers of agro-ecological services in the U.S. The plant growth of these rangelands helps determine the amount of forage available for our livestock and for wildlife, as well as information about fire likelihood and restoration opportunities. However, every spring, ranchers and other rangeland managers face the same difficult challenge —tryingLearning From the Past and Planning for the Future: Experience-Driven Insight Into Managing for Ecosystem Transformations Induced by Drought and Wildfire
Drought and wildfire pose enormous threats to the integrity of natural resources that land managers are charged with protecting. Recent observations and modeling forecasts indicate that these stressors will likely produce catastrophic ecosystem transformations, or abrupt changes in the condition of plants, wildlife, and their habitats, in regions across the country in coming decades. In this pDetermining Successful Management and Restoration Strategies for Pinyon-Juniper Communities in the Face of Changing Climate and Wildfire
Pinyon-juniper woodlands are important ecosystems in the western U.S. that provide numerous critical environmental, economic, and cultural benefits. For example, pinyon pines are a significant cultural resource for multiple Native American Tribes and provide necessary habitat for plants and wildlife (including at risk species, such as the pinyon-jay). Despite their importance, stress put on pinyonImplementing a Grassland Productivity Forecast Tool for the U.S. Southwest
Rangeland systems are some of our nation’s largest providers of agro-ecological services, sustaining plant productivity that is highly variable across seasons and years. Although the ability to predict the upcoming growing season’s rangeland productivity would have enormous economic and management value – such as for making decisions about cattle stocking rates, fire, restoration, and wildlife – t...Long-Term Vegetation Change on the Colorado Plateau
The Colorado Plateau, centered around the four corners area of the Southwest, and includes much of Arizona, Utah, Colorado, and New Mexico, is a large and important component of U.S. drylands. This important home to mountains, desert basins, dramatic canyons, arid woodlands, and grasslands is also one of North America’s most rapidly warming hot spots, with rates of warming of up to 2-3° C within...Novel multi‐scale synthesis of nitrogen fixation rates and drivers across the terrestrial biosphere
Biological nitrogen fixation (BNF) is a critical biogeochemical process that converts inert atmospheric N2 gas into biologically usable forms of the essential nutrient nitrogen. A variety of free-living and symbiotic organisms carry out BNF, and in most regions worldwide, BNF is the largest source of nitrogen that fuels terrestrial ecosystems. As a result, BNF has far reaching effects on ecosystemSouthwest Energy Exploration, Development, and Reclamation (SWEDR)
Deserts of the southwestern US are replete with oil and gas deposits as well as sites for solar, wind, and geothermal energy production. In the past, many of these resources have been too expensive to develop, but increased demand and new technologies have led to an increase in exploration and development. However, desert ecosystems generally have low resilience to disturbance. More frequent... - Data
Meteorological measurements from five weather stations in Grand and San Juan Counties in southeastern Utah (ver. 2.0, April 2023)
These data were compiled from six automated weather stations that together provide several points in the meteorological record across a latitudinal and elevational gradient in southeastern Utah. Recorded data from these weather stations are associated with several different studies. Meteorological data are important for quantifying short term weather events as well as longer term climate trends. SEcological property data and experimental lab incubation results from a long-term nitrogen deposition simulation experiment in three semi-arid grasslands, Arches National Park, Utah, USA, 2013-2019
These data were compiled to understand the responses of dryland ecosystem properties to long-term simulated atmospheric nitrogen deposition. Objective(s) of our study were to uncover any changes in soil biogeochemistry and ecosystem properties to long-term nitrogen amendments. These data represent ecosystem property data compiled over the nine-year history of a nitrogen deposition simulation experData and software code from two long-term experiments (1996-2011 and 2005-2018) at three sites on the Colorado Plateau of North America
These data were compiled to examine how climate change affects biocrust recovery from both physical and climate-induced disturbance. Objective(s) of our study were to uncover the trajectory of biological soil crust communities and soil stability following distrubance and under warming. These data represent biological soil crust surveys under 5 treatments at three sites. These data were collected aCO2 concentrations and microbial biomass data derived from incubation experiments on soils collected at Arches National Park in 2017 and 2018
These data were compiled to improve our understanding of how water, carbon (C), nitrogen (N), and phosphorus (P) interact to regulate below ground carbon cycling. Objective(s) of our study were to evaluate how soil heterotrophic carbon cycling responded to inputs of water, C, N, and P individually and interactively on the Colorado Plateau. These data represent soil microbial and CO2 respiration reRiparian vegetation, topography, and ground cover constituents along the Upper Colorado River near Moab, UT (2010-2017) (ver. 1.1, Jan 2023)
The data release presents observations of riparian vegetation, topography, and ground cover in two river reaches of the Upper Colorado River within a river segment extending 208 river kilometers (rkm), from near the Colorado/Utah border to the confluence of the Green River. Methods included field observations and analysis of the plant community five times over eight years in the fall of 2010, 2012Erosion and Rehabilitation Data, Bandelier National Monument, New Mexico, USA
These data were compiled for a restoration experiment testing the regenerative and functional response of biocrust inoculum reintroduced to a field setting. Regenerative traits measured included measurements of biocrust cover, chlorophyll content, and the roughness of the soil surface. Functional traits measured included nutrient cycling and soil stability. Additionally, these data were compiled f - Multimedia
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Filter Total Items: 150
Time, climate, and soil settings set the course for reclamation outcomes following dryland energy development
Soil attributes, climate, and time since reclamation have important implications for oil and gas reclamation success on drylands. It is uncertain if reclaimed well pads, on highly degraded drylands, can successfully regain ecological function or meet indicator benchmarks for reclamation. Here, our goals were to assess patterns in reclamation outcomes relative to (1) soil attributes, climate, and tAuthorsRandi Corrine Lupardus, Aarin Sengsirirak, Keven Griffen, Anna C Knight, Brandon E McNellis, John B. Bradford, Seth M. Munson, Sasha C. Reed, Miguel L. Villarreal, Michael C. DuniwayStorms and pH of dam releases affect downstream phosphorus cycling in an arid regulated river
Reservoirs often bury phosphorus (P), leading to seasonal or persistent reductions in P supply to downstream rivers. Here we ask if observed variation in the chemistry of dam release waters stimulates downstream sediment P release and biological activity in an arid, oligotrophic system, the Colorado River below Lake Powell, Arizona, USA. We use bottle incubations to simulate a range of observed pHAuthorsBridget Deemer, Robin H. Reibold, Anna Fatta, Jessica R. Corman, Charles Yackulic, Sasha C. ReedCommunity for data integration 2019 project report
The U.S. Geological Survey Community for Data Integration annually supports small projects focusing on data integration for interdisciplinary research, innovative data management, and demonstration of new technologies. This report provides a summary of the 14 projects supported in fiscal year 2019 and outlines their goals, activities, and accomplishments. Proposals in 2019 were encouraged to addreAuthorsAmanda N. Liford, Caitlin M. Andrews, Aparna Bamzai, Joseph A. Bard, David S. Blehert, John B. Bradford, Wesley M. Daniel, Sara L. Caldwell Eldridge, Frank Engel, Jason A. Ferrante, Amy K. Gilmer, Margaret E. Hunter, Jeanne M. Jones, Benjamin Letcher, Frances L. Lightsom, Richard R. McDonald, Leah E. Morgan, Sasha C. Reed, Leslie HsuByEcosystems Mission Area, Water Resources Mission Area, Science Analytics and Synthesis (SAS) Program, Science Synthesis, Analysis and Research Program, Volcano Hazards Program, Community for Data Integration (CDI), Geology, Geophysics, and Geochemistry Science Center, Geosciences and Environmental Change Science Center, National Wildlife Health Center, Oklahoma-Texas Water Science Center, Southwest Biological Science Center, Volcano Science Center, Western Geographic Science Center, Wetland and Aquatic Research Center , Woods Hole Coastal and Marine Science Center, Science Data ManagementLong-term relationships between seed bank communities and wildfire across four North American desert sites
It is well documented that the recovery of dryland plant communities following wildfire can be variable, and that legacies of fire can have long-lasting effects on aboveground plant communities. However, our understanding of the degree to which dryland soil seed banks, or the viable seeds in situ, are impacted by fire and their subsequent postfire succession remains extremely poor. To address thisAuthorsRachel K Hosna, Sasha C. Reed, Akasha M. FaistDroughting a megadrought: Ecological consequences of a decade of experimental drought atop aridification on the Colorado Plateau
Global dryland vegetation communities will likely change as ongoing drought conditions shift regional climates towards a more arid future. Additional aridification of drylands can impact plant and ground cover, biogeochemical cycles, and plant-soil feedbacks, yet how and when these crucial ecosystem components will respond to drought intensification requires further investigation. Using a long-terAuthorsRebecca A Finger-Higgens, Tara Boyce Belnap Bishop, Jayne Belnap, Erika L. Geiger, Edmund E. Grote, David Hoover, Sasha C. Reed, Michael C. DuniwaySoil surface treatments and precipitation timing determine seedling development across southwestern US restoration sites
Restoration in dryland ecosystems often has poor success due to low and variable water availability, degraded soil conditions, and slow plant community recovery rates. Restoration treatments can mitigate these constraints but, because treatments and subsequent monitoring are typically limited in space and time, our understanding of their applicability across broader environmental gradients remainsAuthorsHannah Lucia Farrell, Seth M. Munson, Bradley J. Butterfield, Michael C. Duniway, Aksasha M Faist, Elise S Gornish, Caroline Havrilla, Loralee Larios, Sasha C. Reed, Helen I Rowe, Katherine M. Laushman, Molly L. McCormickGrazing and ecosystem service delivery in global drylands
Grazing represents the most extensive use of land worldwide. Yet its impacts on ecosystem services remain uncertain because pervasive interactions between grazing pressure, climate, soil properties, and biodiversity may occur but have never been addressed simultaneously. Using a standardized survey at 98 sites across six continents, we show that interactions between grazing pressure, climate, soilAuthorsFernando T. Maestre, Yoann Le Bagousse-Pinguet, Manuel Delgado-Baquerizo, David J. Eldridge, Hugo Saiz, Miguel Berdugo, Beatriz Gozalo, Victoria Ochoa, Emilio Guirado, Miguel García-Gómez, Enrique Valencia, Juan J. Gaitán, Sergio Asensio, Betty J. Mendoza, César Plaza, Paloma Díaz-Martínez, Ana Rey, Hang-Wei Hu, Ji-Zheng He, Jun-Tao Wang, Anika Lehmann, Matthias C. Rillig, Simone Cesarz, Nico Eisenhauer, Jaime Martínez-Valderrama, Eduardo Moreno-Jiménez, Osvaldo E. Sala, Mehdi Abedi, Negar Ahmadian, Concepción L. Alados, Valeria Aramayo, Fateh Amghar, Tulio Arredondo, Rodrigo J. Ahumada, Khadijeh Bahalkeh, Farah Ben Salem, Niels Blaum, Bazartseren Boldgiv, Matthew A. Bowker, Donaldo Bran, Chongfeng Bu, Rafaella Canessa, Andrea P. Castillo-Monroy, Helena Castro, Ignacio Castro, Patricio Castro-Quezada, Roukaya Chibani, Abel A. Conceição, Courtney M. Currier, Anthony Darrouzet-Nardi, Balázs Deák, David A. Donoso, Andrew J. Dougill, Jorge Durán, Batdelger Erdenetsetseg, Carlos I. Espinosa, Alex Fajardo, Mohammad Farzam, Daniela Ferrante, Anke S. K. Frank, Lauchlan H. Fraser, Laureano A. Gherardi, Aaron C. Greenville, Carlos A. Guerra, Elizabeth Gusmán-Montalvan, Rosa M. Hernández-Hernández, Norbert Hölzel, Elisabeth Huber-Sannwald, Frederic M. Hughes, Oswaldo Jadán-Maza, Florian Jeltsch, Anke Jentsch, Kudzai F. Kaseke, Melanie Köbel, Jessica E. Koopman, Cintia V. Leder, Anja Linstädter, Peter C. le Roux, Xinkai Li, Pierre Liancourt, Jushan Liu, Michelle A. Louw, Gillian Maggs-Kölling, Thulani P. Makhalanyane, Oumarou Malam Issa, Antonio J. Manzaneda, Eugene Marais, Juan P. Mora, Gerardo Moreno, Seth M. Munson, Alice Nunes, Gabriel Oliva, Gastón R. Oñatibia, Guadalupe Peter, Marco O. D. Pivari, Yolanda Pueyo, R. Emiliano Quiroga, Soroor Rahmanian, Sasha C. Reed, Pedro J. Rey, Benoit Richard, Alexandra Rodríguez, Víctor Rolo, Juan G. Rubalcaba, Jan C. Puppert, Ayman Salah, Max A. Schuchardt, Sedona Spann, Ilan Stavi, Colton R. A. Stephens, Anthony M. Swemmer, Alberto L. Teixido, Andrew D. Thomas, Heather L. Throop, Katja Tielbörger, Samantha K. Travers, James Val, Orsolya Valkó, Liesbeth van den Brink, Sergio Velasco Ayuso, Frederike Velbert, Wanyoike Wamiti, Deli Wang, Lixin Wang, Glenda M. Wardle, Laura Yahdjian, Eli Zaady, Yuanming Zhang, Xiaobing Zhou, Brajesh K. Singh, Nicolas GrossExploring the role of cryptic nitrogen fixers in terrestrial ecosystems: A frontier in nitrogen cycling research
Biological nitrogen fixation represents the largest natural flux of new nitrogen (N) into terrestrial ecosystems, providing a critical N source to support net primary productivity of both natural and agricultural systems. When they are common, symbiotic associations between plants and bacteria can add more than 100 kg N ha−1 y−1 to ecosystems. Yet, these associations are uncommon in many terrestriAuthorsCory Cleveland, Carla R. G. Reis, Steven Perakis, Katherine A Dynarski, Sarah Batterman, Timothy Crews, Maga Gei, Michael Gundale, Duncan Menge, Mark Peoples, Sasha C. Reed, Verity Salmon, Fiona M. Soper, Benton Taylor, Monica Turner, Nina WurzburgerPatterns and controls of foliar nutrient stoichiometry and flexibility across United States forests
Plant element stoichiometry and stoichiometric flexibility strongly regulate ecosystem responses to global change. Here, we tested three potential mechanistic drivers (climate, soil nutrients, and plant taxonomy) of both using paired foliar and soil nutrient data from terrestrial forested National Ecological Observatory Network sites across the USA. We found that broad patterns of foliar nitrogenAuthorsKatherine A Kynarski, Fiona M. Soper, Sasha C. Reed, William R Wieder, Cory C ClevelandUnderstory plant communities show resistance to drought, hurricanes, and experimental warming in a wet tropical forest
Global climate change has led to rising temperatures and to more frequent and intense climatic events, such as storms and droughts. Changes in climate and disturbance regimes can have non-additive effects on plant communities and result in complicated legacies we have yet to understand. This is especially true for tropical forests, which play a significant role in regulating global climate. We useAuthorsAura M. Alonso-Rodríguez, Tana E. Wood, Jamarys Torres-Díaz, Molly A. Cavaleri, Sasha C. Reed, Benedicte BachelotSoutheast Utah Group climate and drought adaptation report: Exposure and perennial grass sensitivity
National Park Service (NPS) managers face growing challenges resulting from the effects of climate change. In particular, as temperatures rise in coming decades, natural resource management in the western United States must cope with expectations for elevated severity and frequency of droughts. These challenges are particularly pronounced for vegetation managers in dryland environments. DevelopingAuthorsJohn B. Bradford, Caroline Havrilla, Jessica A. Hartsell, Daniel Rodolphe Schlaepfer, Molly L. McCormick, Seth M. Munson, Charles Yackulic, Terry T Fisk, David Thoma, Dusty Perkins, Dana L. Witwicki, Matt VanScoyoc, Michael C. Duniway, Sasha C. ReedWhat is a biocrust? A refined, contemporary definition for a broadening research community
Studies of biological soil crusts (biocrusts) have proliferated over the last few decades. The biocrust literature has broadened, with more studies assessing and describing the function of a variety of biocrust communities in a broad range of biomes and habitats and across a large spectrum of disciplines, and also by the incorporation of biocrusts into global perspectives and biogeochemical modelsAuthorsBettina Weber, Jayne Belnap, Burkhard Büdel, Anita J. Antoninka, Nichole N. Barger, V Bala Chaudhary, Anthony Darrouzet-Nardi, David J. Eldridge, Akasha M. Faist, Scott Ferrenberg, Caroline Havrilla, Elisabeth Huber-Sannwald, Oumarou Malam Issa, Fernando T. Maestre, Sasha C. Reed, Emilio Rodriguez-Caballero, Colin L Tucker, Kristina E Young, Yuanming Zhang, Yunge Zhao, Xiaobing Zhou, Matthew A. BowkerNon-USGS Publications**
Sullivan, B.W., Alvarez-Clare, S., Castle, S.C., Porder, S., Reed, S.C., Schreeg, L., Townsend, A.R. and Cleveland, C.C., 2014. Assessing nutrient limitation in complex forested ecosystems: alternatives to large‐scale fertilization experiments. Ecology, 95(3), pp.668-681.Cleveland, C.C., Houlton, B.Z., Smith, W.K., Marklein, A.R., Reed, S.C., Parton, W., Del Grosso, S.J. and Running, S.W., 2013. Patterns of new versus recycled primary production in the terrestrial biosphere. Proceedings of the National Academy of Sciences, 110(31), pp.12733-12737.Reed, S.C., Cleveland, C.C. and Townsend, A.R., 2013. Relationships among phosphorus, molybdenum and free-living nitrogen fixation in tropical rain forests: results from observational and experimental analyses. Biogeochemistry, 114(1-3), pp.135-147.Wickings, K., Grandy, A.S., Reed, S.C. and Cleveland, C.C., 2012. The origin of litter chemical complexity during decomposition. Ecology Letters, 15(10), pp.1180-1188.Reed, S.C., 2008. Scaling from molecules to ecosystems: controls over free-living nitrogen fixation in terrestrial ecosystems. ProQuest.Sattin, S.R., Cleveland, C.C., Hood, E., Reed, S.C., King, A.J., Schmidt, S.K., Robeson, M.S., Ascarrunz, N. and Nemergut, D.R., 2009. Functional shifts in unvegetated, perhumid, recently-deglaciated soils do not correlate with shifts in soil bacterial community composition. The Journal of Microbiology, 47(6), pp.673-681.Costello, E.K., Halloy, S.R., Reed, S.C., Sowell, P. and Schmidt, S.K., 2009. Fumarole-supported islands of biodiversity within a hyperarid, high-elevation landscape on Socompa Volcano, Puna de Atacama, Andes. Applied and Environmental Microbiology, 75(3), pp.735-747.Freeman, K.R., Pescador, M.Y., Reed, S.C., Costello, E.K., Robeson, M.S. and Schmidt, S.K., 2009. Soil CO2 flux and photoautotrophic community composition in high‐elevation,‘barren’soil. Environmental Microbiology, 11(3), pp.674-686.Schmidt, S.K., Reed, S.C., Nemergut, D.R., Grandy, A.S., Cleveland, C.C., Weintraub, M.N., Hill, A.W., Costello, E.K., Meyer, A.F., Neff, J.C. and Martin, A.M., 2008. The earliest stages of ecosystem succession in high-elevation (5000 metres above sea level), recently deglaciated soils. Proceedings of the Royal Society of London B: Biological Sciences, 275(1653), pp.2793-2802.Reed, S.C., Cleveland, C.C. and Townsend, A.R., 2008. Tree species control rates of free-living nitrogen fixation in a tropical rain forest. Ecology, 89(10), pp.2924-2934.Reed, S.C., Cleveland, C.C. and Townsend, A.R., 2007. Controls over leaf litter and soil nitrogen fixation in two lowland tropical rain forests. Biotropica, 39(5), pp.585-592.Schmidt, S.K., Costello, E.K., Nemergut, D.R., Cleveland, C.C., Reed, S.C., Weintraub, M.N., Meyer, A.F. and Martin, A.M., 2007. Biogeochemical consequences of rapid microbial turnover and seasonal succession in soil. Ecology, 88(6), pp.1379-1385.Reed, S.C., Seastedt, T.R., Mann, C.M., Suding, K.N., Townsend, A.R. and Cherwin, K.L., 2007. Phosphorus fertilization stimulates nitrogen fixation and increases inorganic nitrogen concentrations in a restored prairie. Applied Soil Ecology, 36(2), pp.238-242.Cleveland, C.C., Reed, S.C. and Townsend, A.R., 2006. Nutrient regulation of organic matter decomposition in a tropical rain forest. Ecology, 87(2), pp.492-503.Bowker, M.A., Reed, S.C., Belnap, J. and Phillips, S.L., 2002. Temporal variation in community composition, pigmentation, and Fv/Fm of desert cyanobacterial soil crusts. Microbial Ecology, 43(1), pp.13-25.Reed, S.C., Capitosti, G.J., Zhu, Z. and Modarelli, D.A., 2001. Photochemical generation and matrix-isolation detection of dimethylvinylidene. The Journal of Organic Chemistry, 66(1), pp.287-299.Reed, S.C. and Modarelli, D.A., 1996. Conformational effects on the excited state 1, 2-hydrogen migration in alkyldiazomethanes. Tetrahedron Letters, 37(40), pp.7209-7212.**Disclaimer: The views expressed in Non-USGS publications are those of the author and do not represent the views of the USGS, Department of the Interior, or the U.S. Government.
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